Cretornis Temporal range: Late Cretaceous, | |
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Partial wing of Cretornis hlavaci | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Nyctosauromorpha |
Genus: | † Cretornis Frič, 1881 |
Type species | |
†Cretornis hlavaci | |
Synonyms | |
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Cretornis is a pterosaur genus from the late Cretaceous period (Turonian stage) of what is now the Jizera Formation in the Czech Republic, dating to about 92 million years ago. It only contains a single species, Cretornis hlavaci.
The fossils were discovered in 1880 by workers at a sandstone quarry in Zářecká Lhota near the town of Choceň, who were getting gravel to repair a local road. A certain Mrs. Tomková, a croupier from Choceň, then alerted František Hlaváč, a Choceň pharmacist and fossil collector, to the find. He recognised exceptionality of that one, secured the rest of the fossils and then sent them to naturalist Professor Antonín Frič in Prague. In 1881, Antonín Frič identified it as a prehistoric toothed bird the size of a recent swan and named it as the type species Cretornis Hlaváči. [1] The generic name is derived from Latin creta, "chalk", in reference to the Cretaceous, and Greek ὄρνις, ornis, "bird", as Frič originally thought that the fossil bones belonged to some kind of ancient toothed bird (similar to the genus Ichthyornis ). The specific name honors Hlaváč. [2]
Today, the holotype fossils of this pterosaur are stored in the National Museum in Prague. There is also unconfirmed information that similar bones (perhaps also pterosaur bones) were found in other places around the quarry before and during this discovery, but later they were apparently lost.
Afterwards it was realized that the find represented some pterosaur. The name was incorrectly emended by Richard Lydekker into Ornithochirus hlavatschi Fritsch 1881, in 1888. [3] As scientific species names are not allowed to contain diacritical signs, the specific name had to be transcribed. Lydekker chose to write it as if it, and Frič himself, were German, as Bohemia at the time was under a strong German cultural influence. Frič, an ethnic Czech, in 1905 ultimately used the correct transcription Ornithocheirus hlavaci. [4] [5]
Since 1905, the taxon was typically considered a nomen dubium . [6] In 2010 however, Russian paleontologist Alexandr Averianov concluded that it should be possible to determine unique traits. [7] In 2015 he and Czech paleontologist Boris Ekrt published a new description, concluding that Cretornis was a valid taxon. [8]
Cretornis is known from the holotype, presently preserved in the collection of the Národní muzeum (National Museum) in Prague as "Object 10". It was found in what Frič called the Mittlere Iserschichten, today known as the Jizera Formation, dating from the Turonian. It is a partial skeleton lacking the skull. It contains a complete humerus (upper arm bone), an ulna, radius, wrist and two phalanges of the wing finger. [7]
Based on comparison with its relatives, the wingspan of C. hlavaci has by Averianov been estimated at 1.5 to 1.6 meters (4.9 to 5.2 ft). [8] The humerus is 76 millimeters (3 in) in length. [7]
In 2015, Averianov indicated a single autapomorphy, unique derived trait: the distal part of the humerus has a diamond-shaped cross-section. [8]
Cretornis was classified as a species in the genus Ornithocheirus in the nineteenth century. In 1997 Coralia-Maria Jianu suggested it belonged to the Pteranodontidae. [9] In 2010, Averianov thought is it was a member of the Azhdarchidae. More detailed comparisons of the wing bones led Averianov to conclude in 2015 that they belonged to a non-azhdarchid azhdarchoid, probably a member of the group Neoazhdarchia with which it has many features in common. [7] [8] It is probably more advanced than the Thalassodromidae, and shares with Montanazhdarcho the trait that the distal part of the ulna has a joint surface that is placed more proximal than the tubercle. [8] In 2018, paleontologist Nicholas Longrich and colleagues had recovered Cretornis within the family Nyctosauridae, in a more derived position than Alamodactylus and Volgadraco , this contradicts its position within the Azhdarchoidea. [10] The cladogram of their analysis is shown below:
However, subsequent phylogenetic analyses placed Cretornis as a pteranodontian outside Nyctosauridae. [11] [12]
Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids are mainly known for including some of the largest flying animals discovered, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. Previously it was thought that by the end of the Cretaceous, most pterosaur families except for the Azhdarchidae disappeared from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. In several analyses, some taxa such as Navajodactylus, Bakonydraco and Montanazhdarcho were moved from Azhdarchidae to other clades.
Ornithocheirus is a pterosaur genus known from fragmentary fossil remains uncovered from sediments in the United Kingdom and possibly Morocco.
Albisaurus was once thought to be a genus of dinosaur, but is now thought to be a non-dinosaurian archosaur. It was first described by Antonin Fritsch, a Czech palaeontologist, in 1893, but the remains are sparse. The validity of the species cannot be proven based on the fossil remains, and it is usually marked as a nomen dubium. It lived during the Turonian-Santonian stages of the Cretaceous period.
Ponerosteus is a dubious genus of extinct archosauromorph from the Late Cretaceous (Cenomanian-aged) Korycanar Formation of the Czech Republic that was initially identified as a species of the dinosaur Iguanodon.
The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head. The spectacularly-crested Nyctosaurus is sometimes included in this family, though usually placed in its own family, the Nyctosauridae.
Bennettazhia is a genus of tapejaromorph pterosaur which lived during the Albian stage of the Early Cretaceous from what is now the Hudspeth Formation of the state of Oregon in the United States. Although originally identified as a species of the pteranodontoid pterosaur Pteranodon, Bennettazhia is now thought to have been a different animal. The type and only species is B. oregonensis.
Ornithostoma is a genus of pterodactyloid pterosaur that lived during the Early Cretaceous period of Europe, around 110 million years ago. Ornithostoma was once thought to have been a senior synonym of the pteranodontid Pteranodon due to its toothless anatomy and prior naming.
Eoazhdarcho is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is Eoazhdarcho liaoxiensis. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China.
Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.
Lonchodectes was a genus of lonchodectid pterosaur from several formations dating to the Turonian of England, mostly in the area around Kent. The species belonging to it had been assigned to Ornithocheirus until David Unwin's work of the 1990s and 2000s. Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as Amblydectes, Anhanguera, Coloborhynchus, and Ornithocheirus.
"Palaeornis" cliftii is a pterosaur species known from parts of a single humerus found in the early Cretaceous (Valanginian) of the upper Tunbridge Wells Sand Formation, England.
Ornithocheiridae is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods, around 140 to 90 million years ago.
Volgadraco is a genus of pteranodontian pterosaur from the Upper Cretaceous of European Russia.
The Priesener Formation is a Coniacian geologic formation in the Czech Republic. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation.
Nyctosauridae is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly other continents including South America. It was named in 1889 by Henry Alleyne Nicholson and Richard Lydekker.
"Ornithocheirus" buenzeli is a pterosaur species known from parts of a humerus and part of a lower jaw found in late Cretaceous period rocks of the Grünbach Formation, Austria. While it has traditionally been classified in the genus Ornithocheirus, it is more likely an azhdarchid, though due to the fragmentary nature of known fossil remains, it is considered a nomen dubium.
Lonchodraco is a genus of lonchodraconid pterodactyloid pterosaur from the Late Cretaceous of southern England. The genus includes species that were previously assigned to other genera.
Serradraco is a genus of Early Cretaceous pterodactyloid pterosaur from the Valanginian aged Tunbridge Wells Sand Formation in England. Named by Rigal et al. in 2018 with the description of a second specimen, it contains a single species, S. sagittirostris, which was formerly considered a species of Lonchodectes, L. sagittirostris. In 2020, Averianov suggested it did not belong in Lonchodectidae.
Iserosaurus is an extinct genus of sea turtle from the Late Cretaceous of Czech Republic.
The Jizera Formation is a geologic unit of Late Cretaceous (Turonian) age, located in the Czech Republic. It is a unit of the Bohemian Cretaceous Basin. It consists of fluvial to shallow marine sediments and pterosaur fossils are among the remains found in the Jizera Formation. The type locality of the Jizera Formation is a quarry in Zářecká Lhota.