Vesperopterylus Temporal range: Aptian, | |
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Variation in anurognathid jaw shape, notice Vesperopterylus (D) | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Family: | † Anurognathidae |
Subfamily: | † Anurognathinae |
Genus: | † Vesperopterylus Lü et al., 2017 |
Type species | |
†Vesperopterylus lamadongensis Lü et al., 2017 |
Vesperopterylus (meaning "dusk wing") is a genus of anurognathid pterosaur from the Early Cretaceous Jiufotang Formation of China, the geologically youngest member of its group. Notably, Vesperopterylus appears to have a reversed first toe, which would have been suited for gripping; it was likely arboreal, climbing or clinging to tree branches with curved, sharp claws. It also has a relatively short tail, in contrast with its tailless ( Jeholopterus ) and long-tailed ( Dendrorhynchoides ) relatives. It was first described and named by Lü Junchang et al. While the original spelling of the name was Versperopterylus, this was a typo, and was emended by the authors in accordance with the International Code of Zoological Nomenclature. [1]
In 2021, a phylogenetic analysis conducted by Xuefang Wei and colleagues recovered Vesperopterylus within the subfamily Anurognathinae, a subfamily within the family Anurognathidae. Within the subfamily, Vesperopterylus was recovered in a derived position, sister taxon to Anurognathus . [2] Below is a cladogram representing their phylogenetic analysis:
Jeholopterus was a small anurognathid pterosaur known from the Middle to Late Jurassic Daohugou Beds of the Tiaojishan Formation of Inner Mongolia, China, and possibly the Early Cretaceous Sinuiju Formation of North Korea.
Anurognathus is an extinct genus of small pterosaur from the Late Jurassic Altmühltal Formation of Germany.
Batrachognathus is an extinct genus of anurognathid pterosaur from the Late Jurassic Karabastau Formation of the central Asian republic of Kazakhstan. The genus was named in 1948 by the Russian paleontologist Anatoly Nicolaevich Ryabinin. The type species is Batrachognathus volans. The genus name is derived from Greek batrakhos, "frog" and gnathos, "jaw", in reference to the short wide head. The specific epithet means "flying" in Latin.
Anurognathidae is a family of small, short-tailed pterosaurs that lived in Europe, Asia, and possibly North America during the Jurassic and Cretaceous periods. Five genera are known: Anurognathus, from the Late Jurassic of Germany; Jeholopterus, from the Middle to Late Jurassic of China; Dendrorhynchoides, from the Middle Jurassic of China; Batrachognathus, from the Late Jurassic of Kazakhstan; and Vesperopterylus, from the Early Cretaceous of China. Bennett (2007) suggested that the holotype of Mesadactylus, BYU 2024, a synsacrum, belonged to an anurognathid, though this affinity has been questioned by other authors. Mesadactylus is from the Late Jurassic Morrison Formation of the United States. Indeterminate anurognathid remains have also been reported from the Middle Jurassic Bakhar Svita of Mongolia and the Early Cretaceous of North Korea.
Noripterus is a genus of dsungaripterid pterodactyloid pterosaur from Lower Cretaceous-age Lianmuqin Formation in the Junggar Basin of Xinjiang, China. It was first named by Yang Zhongjian in 1973. Additional fossil remains have been recovered from Tsagaantsav Svita, Mongolia.
Chaoyangopterus is a genus of chaoyangopterid pterosaur known from a partial skeleton found in Liaoning, China. Chaoyangopterus was found in rocks dating back to the Aptian-age Lower Cretaceous Jiufotang Formation of Dapingfang, Chaoyang.
Dendrorhynchoides was a genus of anurognathid pterosaur containing only the holotype species D. curvidentatus that is known from the Middle Jurassic Tiaojishan Formation of Qinglong, northern Hebei Province, China.
Feilongus is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Barremian–Aptian-age Lower Cretaceous Yixian Formation of Beipiao, Liaoning, China.
Jidapterus is a genus of chaoyangopterid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. The genus was in 2003 named by Dong Zhiming, Sun Yue-Wu and Wu Shao-Yuan. The type species is Jidapterus edentus. The genus name is derived from Jílín Dàxué or "Jilin University" and a Latinized Greek pteron, "wing". The specific name means "toothless" in Latin.
Azhdarchoidea is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, Tendaguripterus, that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are Quetzalcoatlus, Hatzegopteryx, and Arambourgiania. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea.
Shenzhoupterus is a genus of chaoyangopterid pterosaur from the Jiufotang Formation of modern-day Liaoning, China. Fossil remains of Shenzhoupterus date back to the Early Cretaceous period, approximately 120 million years ago.
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period. It is known for its exceptionally preserved fossils, including those of plants, insects and vertebrates. It is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks. Previously, the Tiaojishan Formation was grouped together with the underlying Haifanggou Formation as a single "Lanqi Formation." The Tiaojishan Formation forms a key part of the Yanliao Biota assemblage, alongside the Haifanggou Formation.
Rhamphorhynchidae is a group of early pterosaurs named after Rhamphorhynchus, that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley. Members of the group possess no more than 11 pairs of teeth in the rostrum, a deltopectoral crest that is constricted at the base but expanded at the distal end, and a bent phalange on the fifth toe.
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.
Archaeoistiodactylus is an extinct genus of wukongopterid pterosaur from the Middle Jurassic of China.
Boreopteridae is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China.
Chuanqilong is a monospecific genus of basal ankylosaurid dinosaur from the Liaoning Province, China that lived during the Early Cretaceous in what is now the Jiufotang Formation. The type and only species, Chuanqilong chaoyangensis, is known from a nearly complete skeleton with a skull of a juvenile individual. It was described in 2014 by Fenglu Han, Wenjie Zheng, Dongyu Hu, Xing Xu, and Paul M. Barrett. Chuanqilong shows many similarities with Liaoningosaurus and may represent a later ontogenetic stage of the taxon.
Luopterus is an extinct genus of anurognathid pterosaur containing only the holotype species L. mutoudengensis that is known from the Middle Jurassic Tiaojishan Formation of Qinglong, northern Hebei Province, China. It was originally named as a species of Dendrorhynchoides in 2012 but it was moved to the genus Luopterus in 2020. Luopterus was originally thought to be from the Early Cretaceous, with a wingspan that is about 40 centimeters (16 in), making it one of the smallest known pterosaurs.
Sinomacrops is a genus of extinct anurognathid pterosaur from the Middle to Late Jurassic periods of what is now the Daohugou Beds of the Tiaojishan Formation in Mutoudeng, Qinglong County of the Hebei province. The remains of Sinomacrops date back to around 164 to 158 million years ago. The type and only known species is Sinomacrops bondei.
Cascocauda is an extinct genus of anurognathid pterosaur from the Late–Middle Jurassic Tiaojishan Formation of Hebei Province, China. The genus contains a single species, C. rong, known from a complete skeleton belonging to a juvenile individual preserved with extensive soft-tissues, including wing membranes and a dense covering of pycnofibres. Some of these pycnofibres appear to be branched, resembling the feathers of maniraptoran theropod dinosaurs, and suggesting that pterosaur pycnofibres may be closely related to feathers in dinosaurs.