Aralosaurus was a genus of hadrosaurid dinosaur that lived during the Late Cretaceous in what is now Kazakhstan. It is known only by a posterior half of a skull and some post-cranial bones found in the Bostobe Formation in rocks dated from the Upper Santonian-Lower Campanian boundary, at about 83.6 Ma. Only one species is known, Aralosaurus tuberiferus, described by Anatoly Konstantinovich Rozhdestvensky in 1968. The genus name means Aral Sea lizard, because it was found to the northeast of the Aral Sea. The specific epithet tuberiferus means bearing a tuber because the posterior part of the nasal bone rises sharply in front of the orbits like an outgrowth. Aralosaurus was originally reconstituted with a nasal arch similar to that of North American Kritosaurus. For many years, Aralosaurus was thus placed in the clade of the Hadrosaurinae. This classification was invalidated in 2004, following the re-examination of the skull of the animal which allowed to identify in Aralosaurus many typical characters of Lambeosaurinae. In particular, this study revealed that Aralosaurus had a hollow bony structure located far in front of the orbits, which communicated with the respiratory tract. This structure being broken at its base, its shape and size remains undetermined. More recently, Aralosaurus has been identified as the most basal Lambeosaurinae, and placed with its close relative Canardia from the upper Maastrichtian of France in the new clade of Aralosaurini.
Azhdarchidae is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well. Azhdarchids are mainly known for including some of the largest flying animals discovered, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs Azhdarcho, Quetzalcoatlus, and Titanopteryx. They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. Previously it was thought that by the end of the Cretaceous, most pterosaur families except for the Azhdarchidae disappeared from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms.
Azhdarcho is a genus of azhdarchid pterosaur from the late Cretaceous Period of the Bissekty Formation of Uzbekistan, as well as the Zhirkindek Formation of Kazakhstan and possibly also the Ialovachsk Formation of Tajikistan. It is known from fragmentary remains including the distinctive, elongated neck vertebrae that characterizes members of the family Azhdarchidae, a family that includes many giant pterosaurs such as Quetzalcoatlus. The name Azhdarcho comes from the Persian word azhdar (اژدر), a dragon-like creature in Persian mythology. The type species is Azhdarcho lancicollis. The specific epithet lancicollis is derived from the Latin words lancea and collum ("neck").
Arambourgiania is a genus of azhdarchid pterosaur that lived during the Maastrichtian age of the Late Cretaceous, in what is now Jordan. Additional fossil remains from the United States and Morocco have also been found, but their assignment to Arambourgiania is only tentative. The original specimen was discovered in the 1940s by a railway worker near Russeifa, Jordan. After examination by paleontologist Camille Arambourg, a new species was named in 1959, Titanopteryx philadelphiae. The generic name means "titan wing", as the fossil was initially misidentified as a huge wing metacarpal, while the specific name refers to the ancient name of Amman, Philadelphia. The genus "Titanopteryx" would later be problematic, as it had already been taken by a fly. Because of this, paleontologist Lev Nessov in 1989 named a new genus, Arambourgiania, in honor of Arambourg. The new species was now known as Arambourgiania philadelphiae.
Bennettazhia is a genus of tapejaromorph pterosaur which lived during the Albian stage of the Early Cretaceous from what is now the Hudspeth Formation of the state of Oregon in the United States. Although originally identified as a species of the pteranodontoid pterosaur Pteranodon, Bennettazhia is now thought to have been a different animal. The type and only species is B. oregonensis.
Bogolubovia is a genus of pterosaur from the Upper Cretaceous Rybushka Formation of Petrovsk, Saratov Oblast, Russia. It is named for Nikolai Nikolaevich Bogolubov, the paleontologist who discovered the remains in 1914.
Eoazhdarcho is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is Eoazhdarcho liaoxiensis. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China.
Phosphatodraco is a genus of azhdarchid pterosaur that lived during the Late Cretaceous of what is now Morocco. In 2000, a pterosaur specimen consisting of five cervical (neck) vertebrae was discovered in the Ouled Abdoun Phosphatic Basin. The specimen was made the holotype of the new genus and species Phosphatodraco mauritanicus in 2003; the genus name means "dragon from the phosphates", and the specific name refers to the region of Mauretania. Phosphatodraco was the first Late Cretaceous pterosaur known from North Africa, and the second pterosaur genus described from Morocco. It is one of the only known azhdarchids preserving a relatively complete neck, and was one of the last known pterosaurs. Additional cervical vertebrae have since been assigned to the genus, and it has been suggested that fossils of the pterosaur Tethydraco represent wing elements of Phosphatodraco.
Azhdarchoidea is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, Tendaguripterus, that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are Quetzalcoatlus, Hatzegopteryx, and Arambourgiania. Other notable pterosaur families belonging to the Azhdarchoidea are the Tapejaridae, the Thalassodromidae, and the Chaoyangopteridae. This group contains some of the most advanced members of the pterosaur order.
Cretornis is a pterosaur genus from the late Cretaceous period of what is now the Jizera Formation in the Czech Republic, dating to about 92 million years ago. It only contains a single species, Cretornis hlavaci.
Volgadraco is a genus of pteranodontian pterosaur from the Upper Cretaceous of European Russia.
The Bostobe Formation is a geological formation in Qaraghandy & Qyzylorda, Kazakhastan whose strata date back to the Late Cretaceous.
Samrukia is a genus of large Cretaceous pterosaurs known only from a single lower jaw discovered in Kazakhstan. The holotype and only known specimen was collected from the Santonian-Campanian age Bostobynskaya Formation in Kyzylorda District. It was described by Darren Naish, Gareth Dyke, Andrea Cau, François Escuillié, and Pascal Godefroit in 2012, and the type species is named Samrukia nessovi. The species is named after Lev Nessov, a paleontologist, and the genus is named after Samruk, a magical bird of Kazakh folklore.
Aerotitan is a genus of azhdarchid pterosaur that lived during the Maastrichtian age of the Late Cretaceous period in what is now Argentina. Its only remains, which consist of a partial snout, were found in the Allen Formation of the Neuquén Basin in northern Patagonia. This specimen was made the holotype of Aerotitan sudamericanus by paleontologist Fernando Novas and colleagues. The generic name combines the Greek word ἀήρ, meaning "air" and Titan, alluding to the pterosaur's large size. The specific name is a reference to its origin, South America.
Eurazhdarcho is a genus of azhdarchid pterosaur that lived during the Maastrichtian age of the Late Cretaceous period in what is now Romania, about 69 million years ago. Starting in 2009, pterosaur fossil remains were unearthed in a layer of the Sebeș Formation in Lancrăm, southwestern Transylvania by paleontologist Mátyás Vremir. In 2013, he, along with paleontologists Alexander Kellner, Darren Naish, and Gareth Dyke would name the new genus and type species Eurazhdarcho langendorfensis. Its generic name is a combination of Europe and the genus Azhdarcho, while its specific name is in reference to its origin from Langendorf. The holotype specimen of Eurazhdarcho consists of a partial skeleton that includes cervical (neck) vertebrae, metacarpal, and phalanx fragments. The related and much larger azhdarchid Hatzegopteryx has also been found in the same location, indicating a possibility that Eurazhdarcho may have a juvenile of this pterosaur. However, the describers deemed this unlikely as its fossils seem to represent an adult specimen.
The Rybushka Formation is a Campanian geologic formation in the Penza and Saratov Oblasts of European Russia. Pterosaur, fish and invertebrate fossils have been recovered from the formation.
The Zhirkindek Formation is a Late Cretaceous geologic formation in Kazakhstan. The primary lithology is sand with interbeds of clay and silt, and represents a coastal to coastal marine environment. The formation has produced numerous fossils, including Lindholmemydid and Trionychid cryptodires, indeterminate remains of dinosaurs and an indeterminate species of pterosaur Azhdarcho.
Mistralazhdarcho is a genus of azhdarchid pterosaur that lived during the Campanian and Maastrichtian ages of the Late Cretaceous period in what is now France. A rich fossil site was discovered in 1992 by paleontologist Xavier Valentin at Velaux–La Bastide Neuve, in the south of France. Pterosaur fossil remains would be subsequently uncovered in the site. They were found in a layer of the Aix-en-Provence Basin at Velaux–La Bastide Neuve and consist of a partial skeleton that includes the skull. These remains would later be made the holotype specimen of the new genus and type species Mistralazhdarcho maggii, named and described in 2018 by Valentin, along with paleontologists Romain Vullo, Géraldine Garcia, Pascal Godefroit, and Aude Cincotta. The generic name combines the mistral, a typical northern wind from the area, and the related genus Azhdarcho. The specific name honors Jean-Pierre Maggi, the mayor of Velaux, who supported the La Bastide Neuve paleontological project.
Cryodrakon is a genus of azhdarchid pterosaur that lived during the late Campanian age of the Late Cretaceous period in what is now Canada, around 76.7 and 74.3 million years ago. Starting in 1972, fossil remains of large azhdarchid pterosaurs have been reported from Alberta. Paleontologists assigned them to the genus Quetzalcoatlus, given that it was the only known azhdarchid from North America back then and because they had limited information about its actual remains, so they simply could not deduce anything different. In 1992, a partial pterosaur skeleton was uncovered in the Dinosaur Park Formation of Alberta. It was partially described in 1995 by paleontologist Philip J. Currie and colleagues, with a subsequent and more complete description in 2005. But it was not until 2019 that it received a new genus and type species, Cryodrakon boreas, named and described by paleontologists David Hone, Michael Habib, and François Therrien. The partial skeleton was made the holotype specimen of this new pterosaur. Its generic name means "cold dragon" in Ancient Greek, in reference to its Canadian origin, while its specific name refers to the Greek god of the north winds, Boreas. All azhdarchid remains from the Dinosaur Park Formation were subsequently referred to Cryodrakon.
Nipponopterus is a genus of azhdarchid pterosaur that lived during the Turonian and Coniacian ages of the Late Cretaceous period in what is now Japan. Its remains, which consist of a partial cervical (neck) vertebra, were found in the Mifune Group, located in the Kumamoto Prefecture in Kyūshū. It was made the holotype specimen of the genus and type species Nipponopterus mifunensis, named and described by paleontologist Xuanyu Zhou and colleagues in 2024. Nipponopterus represents the first pterosaur to be named from Japan.
