Ctenochasmatids | |
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Ctenochasma elegans specimen from Solnhofen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Order: | † Pterosauria |
Suborder: | † Pterodactyloidea |
Clade: | † Ctenochasmatoidea |
Family: | † Ctenochasmatidae Nopsca, 1928 [1] |
Type species | |
† Ctenochasma roemeri Meyer, 1852 | |
Subgroups | |
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Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods.
The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae, [4] though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. [5]
Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set (Andres, 2021). Anders found that three subfamilies fall within the Ctenochasmatidae: Ctenochasmatinae, Gnathosaurinae and Moganopterinae, while also including several basal genera. [6]
Ctenochasmatidae |
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Ctenochasma is a genus of Late Jurassic ctenochasmatid pterosaur belonging to the suborder Pterodactyloidea. Three species are currently recognized: C. roemeri, C. taqueti, and C. elegans. Their fossilized remains have been found in the Solnhofen Limestone of Bavaria, Germany, the "Purbeck Group" of northeastern Germany, and the Calcaires tâchetés of eastern France.
Eosipterus is an extinct genus of ctenochasmatid pterosaur from the Early Cretaceous period of Liaoning, China. Fossil remains of Eosipterus dated back to the Aptian stage of the Early Cretaceous, 125 million years ago.
Gnathosaurus is a genus of ctenochasmatid pterosaur containing two species: G. subulatus, named in 1833 from the Solnhofen Limestone of Germany, and G. macrurus, known from the Purbeck Limestone of the UK. Its fossil remains dated back to the Late Jurassic period.
Pterodactyloidea is one of the two traditional suborders of pterosaurs, and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal rhamphorhynchoids by their short tails and long wing metacarpals. The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like Nyctosaurus and Tupandactylus. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles.
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.
Gegepterus is a genus of ctenochasmatid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China. Only one species is known, G. changi.
Kepodactylus is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Kimmeridgian-Tithonian-age Upper Jurassic Morrison Formation of Colorado, United States.
Plataleorhynchus is a genus of ctenochasmatid pterodactyloid pterosaur from the Late Jurassic to Early Cretaceous periods of what is now the Purbeck Limestone of Dorset, England.
Ctenochasmatoidea is a group of early pterosaurs within the suborder Pterodactyloidea. Their remains are usually found in what were once coastal or lake environments. They generally had long wings, long necks, and highly specialized teeth.
Dolicorhamphus is an extinct genus of pterosaurs from the Middle Jurassic Taynton Limestone Formation and Fuller's Earth Formations of England. The genus contains two species, D. bucklandii and D. depressirostris.
Elanodactylus is a genus of ctenochasmatid pterodactyloid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China.
Monofenestrata is an unranked group of pterosaurs that includes the family Wukongopteridae and the suborder Pterodactyloidea.
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested Tropeognathus.
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.
Archaeopterodactyloidea is an extinct clade of pterodactyloid pterosaurs that lived from the middle Late Jurassic to the latest Early Cretaceous periods of Africa, Asia, Europe and North America. It was named by Alexander Wilhelm Armin Kellner in 1996 as the group that contains Germanodactylus, Pterodactylus, the Ctenochasmatidae and the Gallodactylidae. In 2003, Kellner defined the clade as a node-based taxon consisting of the last common ancestor of Pterodactylus, Ctenochasma and Gallodactylus and all its descendants. Although phylogenetic analyses that based on David Unwin's 2003 analysis do not recover monophyletic Archaeopterodactyloidea, phylogenetic analyses that based on Kellner's analyses, or the analyses of Brian Andres recover monophyletic Archaeopterodactyloidea at the base of the Pterodactyloidea.
Kryptodrakon is an extinct genus of pterodactyloid pterosaur from the Middle to Late Jurassic with an age of approximately 162.7 million years. It is known from a single type species, Kryptodrakon progenitor. The age of its fossil remains made Kryptodrakon the basalmost and oldest pterodactyloid known to date.
Liaodactylus is a genus of filter-feeding ctenochasmatid pterosaur from the Jurassic of China. The genus contains one species, L. primus, described by Zhou et al. in 2017. As an adaptation to filter-feeding, Liaodactylus had approximately 150 long, comb-like teeth packed closely together. It is both the earliest known ctenochasmatid and the first filter-feeding pterosaur from the Jurassic Yanliao Biota. Later and more specialized ctenochasmatids differ from Liaodactylus in having longer snouts, smaller openings in the skull, and more teeth. Within the Ctenochasmatidae, Liaodactylus was most closely related to the European Ctenochasma.
Tacuadactylus is a genus of ctenochasmatid pterosaur from the Late Jurassic of Uruguay.
Cratonopterus is an extinct genus of ctenochasmatid pterosaurs from the Early Cretaceous Huajiying Formation of China. The genus contains a single species, C. huabei, known from a partial skeleton. Cratonopterus represents the first named pterosaur from the Huajiying Formation.