| Boreopterus Temporal range: Early Cretaceous, | |
|---|---|
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Order: | † Pterosauria |
| Suborder: | † Pterodactyloidea |
| Family: | † Boreopteridae |
| Subfamily: | † Boreopterinae |
| Genus: | † Boreopterus Lü & Ji, 2005 |
| Type species | |
| †Boreopterus cuiae Lü & Ji, 2005 | |
| Other species | |
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| Synonyms | |
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Boreopterus is a genus of boreopterid pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Dalian, Liaoning, China.
Boreopterus was named in 2005 by Lü Junchang and Ji Qiang. The type species is Boreopterus cuiae. The genus name is derived from Greek boreios, "northern" and pteron, "wing". The specific epithet honors Cui Xu.
Boreopterus is based on holotype JZMP-04-07-3, a nearly complete but crushed skeleton and skull. The skull is 235 millimeters long (9.25 inches), low and elongated with a rounded tip. Its wingspan is estimated to have been around 1.45 meters (4.76 feet). Its teeth, especially the anterior nine pairs, are quite large, forming a mesh of sharp teeth at the front of the mouth; the third and fourth teeth from the front are the largest. There are at least 27 teeth in each side of both the upper and lower jaws, which is a large amount. [3]
Lü and Ji initially placed Boreopterus in the family Ornithocheiridae when they described it in 2006, a classification which was supported later that year by David Unwin. [4] However, Lü in 2006 published a cladistic analysis showing Boreopterus to be the sister taxon of Feilongus (together forming the new family Boreopteridae [5] ) in a more basal position than Haopterus . [6]
In 2013, a more comprehensive study of pterosaur relationships supported the close relationship of Boreopterus and Feilongus, as well as their relatively basal status among pterodactyloids. Later that year, Andres & Myers found the boreopterids as the sister group of Cycnorhamphus within the archaeopterodactyloid group Gallodactylidae. [7] However, subsequent analysis have found boreopterids to be indeed members of the Ornithocheiroidea, composed of Boreopterus, Zhenyuanopterus and Guidraco , while Feilongus is a relative of Gnathosaurus . [8] In 2018, Nicholas Longrich and colleagues found the family Boreopteridae to only contain Boreopterus and Zhenyuanopterus, while Guidraco was placed in a more derived position within the clade Anhangueria. They also found the family Lonchodectidae as the sister taxon of Boreopteridae. [9]
Topology 1: Andres et al. (2014). [8]
| Topology 2: Longrich et al. (2018). [9]
|
Pterosaurs like Boreopterus are interpreted by Unwin as soaring animals, like today's albatrosses and frigatebirds. [10] However, it has also been suggested that boreopterids foraged while swimming, trapping small prey with their needle-like teeth, [11] a method similar to that of modern Platanista dolphins.
It has been suggested that the closely related Zhenyuanopterus was merely the adult form of this animal. [11]
Eosipterus is an extinct genus of ctenochasmatid pterosaur from the Early Cretaceous period of Liaoning, China. Fossil remains of Eosipterus dated back to the Aptian stage of the Early Cretaceous, 124.6 million years ago.
The Rhamphorhynchoidea forms one of the two suborders of pterosaurs and represents an evolutionary grade of primitive members of flying reptiles. This suborder is paraphyletic unlike the Pterodactyloidea, which arose from within the Rhamphorhynchoidea as opposed to a more distant common ancestor. Because it is not a completely natural grouping, Rhamphorhynchoidea is not used as a formal group in most scientific literature, though some pterosaur scientists continue to use it as an informal grouping in popular works, such as The Pterosaurs: From Deep Time by David Unwin, and in some formal studies. Rhamphorhynchoids were the first pterosaurs to have appeared, in the late Triassic Period. Unlike their descendants, the pterodactyloids, most rhamphorhynchoids had teeth and long tails, and most species lacked a bony crest, though several are known to have crests formed from soft tissue like keratin. They were generally small, with wingspans rarely exceeding 2.5 meters, though one specimen alluded to by Alexander Stoyanow would be among the largest pterosaurs of all time with a wingspan of 10 meters, comparable to the largest azhdarchids. However, this alleged giant Jurassic pterosaur specimen is not recorded anywhere outside the original Time article. Nearly all rhamphorhynchoids had become extinct by the end of the Jurassic Period, though some anurognathids persisted to the early Cretaceous. The family Wukongopteridae, which shows a mix of rhamphorhynchoid and pterodactyloid features, is known from the Daohugou Beds which are most commonly dated to the Jurassic, but a few studies give a Cretaceous date. Furthermore, remains of a non-pterodactyloid from the Candeleros Formation extend the presence of basal pterosaurs into at least the early Late Cretaceous.
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey.
Beipiaopterus is a genus of ctenochasmatid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of the China.
Eopteranodon is a genus of tapejarid pterosaur from the Aptian-age Lower Cretaceous Yixian Formation of Beipiao City, Liaoning, China. The genus was named in 2005 by paleontologists Lü Junchang and Zhang Xingliao. The type and only species is Eopteranodon lii.
Chaoyangopterus is a genus of chaoyangopterid pterosaur known from a partial skeleton found in Liaoning, China. Chaoyangopterus was found in rocks dating back to the Aptian-age Lower Cretaceous Jiufotang Formation of Dapingfang, Chaoyang.
Eoazhdarcho is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is Eoazhdarcho liaoxiensis. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China.
Feilongus is an extinct genus of ctenochasmatid pterodactyloid pterosaur from the Barremian–Aptian-age Lower Cretaceous Yixian Formation of Beipiao, Liaoning, China.
Haopterus is a genus of pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Its fossil remains dated back 124.6 million years ago.
Liaoxipterus is a genus of pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. The type species is Liaoxipterus brachyognathus. The genus name is derived from the discovery locality Liaoxi and a Latinised Greek pteron, "wing". The specific name is derived from Greek brachys, "short" and gnathos, "jaw".
Nurhachius is a genus of istiodactylid pterodactyloid pterosaur from the Barremian to Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. Its fossil remains date back about 120 million years ago.
Lonchodectidae or Lonchodraconidae is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontia. They are notable for their high, conical tooth sockets and raised alveolar margins.
Elanodactylus is a genus of ctenochasmatid pterodactyloid pterosaur from the Early Cretaceous period of what is now the Yixian Formation of Liaoning, China.
Zhenyuanopterus is a genus of boreopterid pterosaur which is known from Lower Cretaceous Yixian Formation of Liaoning, China. It contains one species, Zhenyuanopterus longirostris, which was first described and named by Lü Junchang.
Rhamphorhynchidae is a group of early pterosaurs named after Rhamphorhynchus, that lived in the Late Jurassic. The family Rhamphorhynchidae was named in 1870 by Harry Govier Seeley. Members of the group possess no more than 11 pairs of teeth in the rostrum, a deltopectoral crest that is constricted at the base but expanded at the distal end, and a bent phalange on the fifth toe.
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of Pterodactylus and Ctenochasma, and all their descendants.
Boreopteridae is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China.
Germanodactylidae is a controversial group of pterosaurs within the suborder Pterodactyloidea. It was first named by Yang Zhongjian in 1964, and given a formal phylogenetic definition in 2014 by Brian Andres, James Clark, and Xu Xing. They defined it as the least inclusive clade containing Germanodactylus cristatus and Normannognathus wellnhoferi, which they considered to be close relatives at the time. However, more recent studies by the same researchers have found that these pterosaurs may be only distantly related.
Moganopterus is an extinct genus of ctenochasmatid pterosaur from the Early Cretaceous of western Liaoning Province, China.
