Spinosaurus is a genus of spinosaurid dinosaur that lived in what now is North Africa during the Cenomanian to upper Turonian stages of the Late Cretaceous period, about 100 to 93.5 million years ago. The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. The original remains were destroyed in World War II, but additional material came to light in the early 21st century. It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The best known species is S. aegyptiacus from Egypt, although a potential second species, S. maroccanus, has been recovered from Morocco. The contemporary spinosaurid genus Sigilmassasaurus has also been synonymized by some authors with S. aegyptiacus, though other researchers propose it to be a distinct taxon. Another possible junior synonym is Oxalaia from the Alcântara Formation in Brazil.
Carcharodontosaurus is a genus of carnivorous theropod dinosaur that lived in North Africa from about 100 to 94 million years ago during the Cenomanian stage of the Late Cretaceous. Two teeth of the genus, now lost, were first described from Algeria by French paleontologists Charles Depéret and Justin Savornin as Megalosaurus saharicus. A partial skeleton was collected by crews of German paleontologist Ernst Stromer during a 1914 expedition to Egypt. Stromer did not report the Egyptian find until 1931, in which he dubbed the novel genus Carcharodontosaurus, making the type species C. saharicus. Unfortunately, this skeleton was destroyed during the Second World War. In 1995 a nearly complete skull of C. saharicus, the first well-preserved specimen to be found in almost a century, was discovered in the Kem Kem Beds of Morocco; it was designated the neotype in 1996. Fossils unearthed from the Echkar Formation of northern Niger were described and named as another species, C. iguidensis, in 2007.
Spinosauridae is a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America and Asia. Their remains have generally been attributed to the Early to Mid Cretaceous.
Deltadromeus is a genus of theropod dinosaur from the Aoufous Formation of Morocco.
Sigilmassasaurus is a controversial genus of spinosaurid dinosaur that lived approximately 100 to 94 million years ago during the Late Cretaceous Period in what is now northern Africa. Named in 1996 by Canadian paleontologist Dale Russell, it contains a single species, Sigilmassasaurus brevicollis. The identity of the genus has been debated by scientists, with some considering its fossils to represent material from the closely related species Spinosaurus aegyptiacus, while others have classified it as a separate taxon, forming the clade Spinosaurini with Spinosaurus as its sister taxon.
Siroccopteryx is an extinct genus of anhanguerid pterodactyloid pterosaur, known from middle Cretaceous sediments in modern-day Morocco. Some researchers, such as David M. Unwin, consider the genus a junior synonym of Coloborhynchus.
The Cambridge Greensand is a geological unit in England whose strata are earliest Cenomanian in age. It lies above the erosive contact between the Gault Formation and the Chalk Group in the vicinity of Cambridgeshire, and technically forms the lowest member bed of the West Melbury Marly Chalk Formation. It is a remanié deposit, containing reworked fossils of late Albian age, including those of dinosaurs and pterosaurs.
Bahariasauridae is a potential family of averostran theropods that might include a handful of African and South American genera, such as Aoniraptor, Bahariasaurus, Deltadromeus, and Gualicho. The placement of these theropods is controversial, with some studies placing them as basal ceratosaurs possibly related to Noasauridae, others classifying them as megaraptorans, basal neovenatorids, or basal coelurosaurs. There is also a possibility the group might not be monophyletic, as a monograph on the vertebrate diversity in the Kem Kem Beds published in 2020 found Bahariasaurus to be nomen dubium. In the same paper Deltadromeus is classified as an noasaurid, a result also recovered by some previous studies. A 2024 analysis found Aoniraptor, Bahariasaurus, Deltadromeus, Elaphrosaurus and Gualicho to form a monophyletic clade at the base of Ceratosauria.
Alanqa is a genus of pterodactyloid pterosaur from the Late Cretaceous period of what is now the Kem Kem Beds of southeastern Morocco. The name Alanqa comes from the Arabic word العنقاءal-‘Anqā’, for a mythical bird of Arabian culture.
Oxalaia is a genus of spinosaurid dinosaur that lived in what is now the Northeast Region of Brazil during the Cenomanian stage of the Late Cretaceous period, sometime between 100.5 and 93.9 million years ago. Its only known fossils were found in 1999 on Cajual Island in the rocks of the Alcântara Formation, which is known for its abundance of fragmentary, isolated fossil specimens. The remains of Oxalaia were described in 2011 by Brazilian palaeontologist Alexander Kellner and colleagues, who assigned the specimens to a new genus containing one species, Oxalaia quilombensis. The species name refers to the Brazilian quilombo settlements. Oxalaia quilombensis is the eighth officially named theropod species from Brazil and the largest carnivorous dinosaur discovered there. It is closely related to the African genus Spinosaurus, and/or may be a junior synonym of this taxon.
Sauroniops is a controversial genus of carnivorous basal carcharodontosaurid theropod dinosaur known from the Late Cretaceous Gara Sbaa Formation, and possibly also the Kem Kem Formation, both of Morocco. The type, and currently only, species is S. platytholus.
Gualicho is a genus of theropod dinosaur. The type species is Gualicho shinyae. It lived in what is now northern Patagonia, on what was then a South American island continent split off from the supercontinent Gondwana. The fossils were found in the Huincul Formation, dating to the late Cenomanian-early Turonian age of the upper Cretaceous Period, around 91 million years ago.
Xericeps is a genus of pterosaur from the Late Cretaceous Kem Kem Beds of southeastern Morocco.
Nizar Ibrahim is a German-Moroccan vertebrate paleontologist and comparative anatomist. He is currently a senior lecturer at the University of Portsmouth. Ibrahim has led several expeditions to Africa's Sahara and is notable for his research on fossil vertebrates from the Kem Kem Group, including pterosaurs, crocodyliforms, and dinosaurs. In recent years, research led by Ibrahim radically changed ideas about the morphology and life habits of one of the largest predatory dinosaurs, Spinosaurus aegyptiacus. Ibrahim also has interests in bioinformatics and contributed to the NSF-funded Phenoscape project. He regularly engages with the public and is a speaker with the National Geographic Speakers Bureau.
Afrotapejara is an extinct genus of tapejarid pterosaur discovered in Morocco. The type species, Afrotapejara zouhri, was named and described in 2020. It was the first tapejarid discovered in Africa and the fourth pterosaur discovered in the Kem Kem Beds.
Apatorhamphus is an extinct genus of azhdarchoid pterosaur from the Kem Kem Group of Morocco. It might have been part of the Chaoyangopteridae. It is only known from a few snout fragments and it likely had a wingspan of between 3–7 metres (9.8–23.0 ft)
Leptostomia is a genus of long-beaked pterosaur from the mid-Cretaceous (?Albian-Cenomanian) of Morocco, North Africa. The type species is L. begaaensis, which was named and described in 2021 from sediments of the Kem Kem Group in Morocco. It was a small animal with a long, slender bill which is thought to have been used to probe sediments for worms and other invertebrates, similar to kiwi birds and curlews. Leptostomia is likely a member of the Azhdarchoidea.
Nicorhynchus is a genus of anhanguerid pterosaur from the Cretaceous period. It contains two species, the type species, N. capito, from the Cambridge Greensand of England, and N. fluviferox from the Kem Kem Group of Morocco. These species were previously assigned to Coloborhynchus.
Oniichthys is an extinct genus of gar in the family Lepisosteidae. It contains a single species, O. falipoui, known from the Late Cretaceous (Cenomanian) of Morocco.
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