Aegyptosuchidae Temporal range: Late Cretaceous: Cenomanian, | |
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Skull of Aegisuchus witmeri | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauria |
Clade: | Pseudosuchia |
Clade: | Crocodylomorpha |
Clade: | Crocodyliformes |
Clade: | Neosuchia |
Clade: | Eusuchia |
Family: | † Aegyptosuchidae Kuhn, 1936 |
Genera | |
Aegyptosuchidae is an extinct family of eusuchian crocodyliforms from the Cretaceous period of Africa. They are characterized by their large size and flat heads. The family includes two genera, Aegyptosuchus and Aegisuchus . [1]
Aegyptosuchidae was originally established as the monotypic family name for Aegyptosuchus peyeri in 1933. Upon the discovery of the fellow aegyptosuchid Aegisuchus witmeri in 2012 by Holliday and Gardner, Aegyptosuchidae was phylogenetically defined as the least inclusive clade containing Aegisuchus witmeri and Aegyptosuchus peyeri, so long as it does not include Alligator mississippiensis (American alligator), Bernissartia fagesii , Crocodylus niloticus (Nile crocodile), Gavialis gangeticus (gharial), Hylaeochampsa vectiana , or Susisuchus anatoceps . [1]
Aegyptosuchidae belongs to the clade Eusuchia, and is proposed to be the sister clade to the crown group Crocodylia, which contains all extant (living) crocodilians. The phylogeny can be shown in the cladogram below: [1]
Gavialidae is a family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two living species, the gharial and the false gharial, both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct Hanyusuchus. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a generalist diet with mature adults preying upon larger vertebrates, such as ungulates.
Allodaposuchus is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages, and possibly the Santonian stage, of the Late Cretaceous. Although generally classified as a non-crocodylian eusuchian crocodylomorph, it is sometimes placed as one of the earliest true crocodylians. Allodaposuchus is one of the most common Late Cretaceous crocodylomorphs from Europe, with fossils known from Romania, Spain, and France.
Boverisuchus is an extinct genus of planocraniid crocodyliforms known from the middle Eocene of Germany and western North America. It was a relatively small crocodyliform with an estimated total length of approximately 2.2–3.6 metres (7.2–11.8 ft).
Argochampsa is an extinct genus of eusuchian crocodylomorph, usually regarded as a gavialoid crocodilian, related to modern gharials. It lived in the Paleocene of Morocco. Described by Hua and Jouve in 2004, the type species is A. krebsi, with the species named for Bernard Krebs. Argochampsa had a long narrow snout, and appears to have been marine in habits.
Aegyptosuchus is an extinct monospecific genus of aegyptosuchid eusuchian crocodyliform. It was found in the Bahariya Formation of Egypt, which dates back to the Cenomanian age of the Late Cretaceous. The type and only species is Aegyptosuchus peyeri.
Eusuchia is a clade of neosuchian crocodylomorphs that first appeared in the Early Cretaceous, which includes modern crocodilians. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-Pg extinction.
Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.
Acynodon is an extinct genus of eusuchian crocodylomorph from the Late Cretaceous, with fossils found throughout Southern Europe.
Hylaeochampsidae is an extinct family of basal eusuchian crocodylomorphs thought to be closely related to the order Crocodylia.
Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudosuchians to survive the K-Pg extinction event.
Planocrania is an extinct genus of eusuchian crocodyliforms from what is now China. Two species are currently known to belong to the genus.
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to the Holocene, although this is disputed.
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial Gavialis gangeticus and the false gharial Tomistoma schlegelii are alive today, with Hanyusuchus having become extinct in the last few centuries.
Brevirostres is a paraphyletic group of crocodilians that included alligatoroids and crocodyloids. Brevirostres are crocodilians with small snouts, and are distinguished from the long-snouted gharials. It is defined phylogenetically as the last common ancestor of Alligator mississippiensis and Crocodylus niloticus and all of its descendants. This classification was based on morphological studies primarily focused on analyzing skeletal traits of living and extinct fossil species, and placed the gharials outside the group due to their unique skull structure, and can be shown in the simplified cladogram below:
Aegisuchus is an extinct monospecific genus of giant, flat-headed crocodyliform within the family Aegyptosuchidae. It was found in the Kem Kem Formation of southeast Morocco, which dates back to the Cenomanian age of the Late Cretaceous epoch. The type species Aegisuchus witmeri was named in 2012 by paleontologists Casey Holliday and Nicholas Gardner, who nicknamed it "Shieldcroc" for the shield-like shape of its skull. A. witmeri is known from a single partial skull including the braincase and skull roof.
Planocraniidae is an extinct family of eusuchian crocodyliforms known from the Paleogene of Asia, Europe and North America. The family was coined by Li in 1976, and contains three genera, Boverisuchus, Duerosuchus and Planocrania. Planocraniids were highly specialized crocodyliforms that were adapted to living on land. They had extensive body armor, long legs, and blunt claws resembling hooves, and are sometimes informally called "hoofed crocodiles".
Allodaposuchidae is an extinct clade of eusuchians that lived in Europe during the Late Cretaceous (Santonian-Maastrichtian).
Lohuecosuchus is an extinct genus of allodaposuchid eusuchian crocodylomorph that lived during the Late Cretaceous in what is now Spain and southern France.
Portugalosuchus is an extinct genus of eusuchian crocodyliform that was possibly a basal crocodylian – if so then it would be the oldest known crocodylian to date. The type species is P. azenhae, described in 2018, and it is known from the Late Cretaceous (Cenomanian)-aged Tentugal Formation in Portugal. A 2021 morphological study recovered Portugalosuchus within Crocodylia as a member of Gavialidae closely related to similar "thoracosaurs", while also noting that it might also possibly be outside of Crocodylia completely. A 2022 tip dating analysis incorporating both morphological and DNA data placed Portugalosuchus outside of Crocodylia, as the sister taxon of the family Allodaposuchidae. A cladogram simplified after that analysis is shown below:
Longirostres is a clade of crocodilians that includes the crocodiles and the gavialids, to the exclusion of the alligatoroids. Defined in 2003 by Harshman et al., Longirostres is a crown group defined phylogenetically as including the last common ancestor of Crocodylus niloticus and Gavialis gangeticus and all of its descendants.