Argochampsa

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Argochampsa
Temporal range: Paleocene,
66–56  Ma [1]
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pseudosuchia
Superorder: Crocodylomorpha
Clade: Eusuchia
Genus: Argochampsa
Hua & Jouve 2004
Type species
Argochampsa krebsi

Argochampsa (meaning "Argo crocodile", in reference to the mythological Argo of Jason) is an extinct genus of eusuchian crocodylomorph, usually regarded as a gavialoid crocodilian, related to modern gharials. It lived in the Paleocene of Morocco. Described by Hua and Jouve in 2004, the type species is A. krebsi, with the species named for Bernard Krebs  [ de ]. Argochampsa had a long narrow snout, and appears to have been marine in habits. [2]

Contents

Description

Argochampsa is based on OCP DEK-GE 1201, a nearly complete skull from the Oulad Abdoun Basin, in the vicinity of Khouribga, Morocco. The skull, 43.3 centimeters long (17.0 in), had a long, narrow snout, marking it as a longirostrine crocodilian; the snout made up about 70% of the skull's length. The premaxillae at the tip of the snout were downturned, and the tip somewhat squared off, with the first few tooth positions in a straight line perpendicular to the long axis of the skull. There were five teeth in each premaxilla and 26 in each maxilla (the main tooth-bearing bone of the upper jaw). The nasal bones were fused, and there were several short diastemas or gaps in the tooth row at the tip of the snout. [2] More recently, material from the lower jaw, neck and back vertebrae, the upper arm, and armor has been recovered. [3]

Hua and Jouve performed a cladistic analysis incorporating their new taxon, and found Argochampsa to be a gavialoid, but outside of the clade Gavialidae. They noted that the snout shape of Argochampsa is unusual among crocodilians, with only pholidosaurids and Terminonaris having similarly shaped snout tips, and suggested that this layout may have facilitated precise occlusion. Argochampsa lived in an environment otherwise dominated by dyrosaurid marine crocodyliforms. [2]

Argochampsa krebsi was included in the study on the phylogenetic relationships of putative fossil gavialoids published by Lee & Yates (2018). The authors considered it most likely that Argochampsa was not a gavialoid, or even a crocodylian, but rather a member of the clade of non-crocodylian eusuchians that also included the genera Eogavialis , Eosuchus , Eothoracosaurus and Thoracosaurus . [4]

Related Research Articles

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<i>Gavialosuchus</i> Extinct genus of reptiles

Gavialosuchus is an extinct genus of gavialid crocodylian from the early Miocene of Europe. Currently only one species is recognized, as a few other species of Gavialosuchus have since been reclassified to other genera.

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<span class="mw-page-title-main">Eusuchia</span> Clade of reptiles

Eusuchia is a clade of crocodylomorphs that first appeared in the Early Cretaceous with Hylaeochampsa. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades died out 47 and 11 million years ago respectively, all living crocodilian species are eusuchians, as are many extinct forms.

<i>Navajosuchus</i> Extinct genus of reptiles

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Eothoracosaurus is an extinct monospecific genus of eusuchian crocodylomorphs found in Eastern United States which existed during the Late Cretaceous period. Eothoracosaurus is considered to belong to an informally named clade called the "thoracosaurs", named after the closely related Thoracosaurus. Thoracosaurs in general were traditionally thought to be related to the modern false gharial, largely because the nasal bones contact the premaxillae, but phylogenetic work starting in the 1990s instead supported affinities within gavialoid exclusive of such forms. Even more recent phylogenetic studies suggest that thoracosaurs might instead be non-crocodilian eusuchians.

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<i>Planocrania</i> Extinct genus of reptiles

Planocrania is an extinct genus of eusuchian crocodyliforms from what is now China. Two species are currently known to belong to the genus.

Phosphatosaurus is an extinct genus of dyrosaurid crocodylomorph. It existed during the early Eocene, with fossils having been found from North Africa in Tunisia and Mali. Named in 1955, Phosphatosaurus is a monotypic genus; the type species is P. gavialoides. A specimen has been discovered from Niger, but it cannot be classified at the species level.

<i>Piscogavialis</i> Extinct genus of reptiles

Piscogavialis is an extinct monospecific genus of gryposuchine gavialid crocodylian. The only species yet known is P. jugaliperforatus. Fossils of Piscogavialis have been found from the Mio-Pliocene Pisco Formation of the Sacaco Basin in southern Peru in 1998, where it coexisted with the much smaller gavialid Sacacosuchus.

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<span class="mw-page-title-main">Gavialoidea</span> Superfamily of large reptiles

Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial Gavialis gangeticus and the false gharial Tomistoma schlegelii are alive today, with Hanyusuchus having become extinct in the last few centuries.

<span class="mw-page-title-main">Planocraniidae</span> Extinct family of reptiles

Planocraniidae is an extinct family of eusuchian crocodyliforms known from the Paleogene of Asia, Europe and North America. The family was coined by Li in 1976, and contains three genera, Boverisuchus, Duerosuchus and Planocrania. Planocraniids were highly specialized crocodyliforms that were adapted to living on land. They had extensive body armor, long legs, and blunt claws resembling hooves, and are sometimes informally called "hoofed crocodiles".

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References

  1. Rio, Jonathan P.; Mannion, Philip D. (6 September 2021). "Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem". PeerJ . 9: e12094. doi: 10.7717/peerj.12094 . PMC   8428266 . PMID   34567843.
  2. 1 2 3 Hua, Stephane; Jouve, Stéphane (2004). "A primitive marine gavialoid from the Paleocene of Morocco" (PDF). Journal of Vertebrate Paleontology. 24 (2): 341–350. doi:10.1671/1104. S2CID   86497272.
  3. Jouve, Stéphane; Iarocheneb, Mohamed; Bouyac, Baâdi; Amaghzaz, Mbarek (2006). "New material of Argochampsa krebsi (Crocodylia: Gavialoidea) from the Lower Paleocene of the Oulad Abdoun Basin (Morocco): phylogenetic implications". Geobios. 39 (6): 817–832. doi:10.1016/j.geobios.2005.07.003.
  4. Michael S. Y. Lee; Adam M. Yates (2018). "Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil record". Proceedings of the Royal Society B: Biological Sciences. 285 (1881): 20181071. doi:10.1098/rspb.2018.1071. PMC   6030529 . PMID   30051855.