Rugosuchus Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauria |
Clade: | Pseudosuchia |
Clade: | Crocodylomorpha |
Clade: | Crocodyliformes |
Family: | † Paralligatoridae |
Genus: | † Rugosuchus Wu et al., 2001 |
Type species | |
†Rugosuchus nonganensis Wu et al., 2001 |
Rugosuchus (meaning "uneven or wrinkled crocodile", in reference to texturing on its upper jaw bones) is an extinct genus of neosuchian crocodyliform [1] from the Cretaceous of China. It is known from most of a skull, a partial postcranial skeleton, and a second partial skeleton including part of the hips. It was described by Xiao-Chun Wu and colleagues in 2001, with R. nonganensis as the type species. At the time of its description, it was the most complete crocodyliform from northeastern China, and only the second known.
Rugosuchus is based on IGV 33, most of a skull. Wu et al. assigned two other specimens to this genus: IGV 31, much of a skeleton lacking a skull and most of the limbs; and IGV 32, three vertebrae, a partial hip, and a fragment of thigh bone. These three specimens had been collected in 1958 by the Petroleum Geological Survey of the Song-Liao Basin, and remained unstudied for many years. The locality is near Fulongquan, Nong'an County, Jilin, in the Song-Liao Basin. The formation is not known for certain, but is probably the Nenjiang Formation. The age of the Nenjiang Formation has been debated, but as of the description of Rugosuchus was thought to be latest Early Cretaceous, based on ostracode, bivalve, and fish fossils. [2] However, most of later studies consider it as late Cretaceous instead. [3]
Skull IGV 33 is thought to have come from an adult, based on the fusion of bones. It is somewhat elongate, being about 28 centimeters (11 in) long but only an estimated 13 centimeters (5 in) at its widest. The maxillae, the main tooth-bearing bones of the upper jaw, had unusual elongate depressions on their sides, nine or ten per maxilla. Other skull bones had heavily textured surfaces, as is seen on other crocodyliforms. The premaxillae at the tip of the snout had five teeth each, and the maxillae 16 or 17, with wide spacing; because the jaws are closed, the teeth of the lower jaw cannot be observed. [2]
Wu et al. noted that their new genus was not an eusuchian because of the form of the palatal bones, but it was probably more closely related to the eusuchians than other extinct crocodyliforms like Goniopholis by having more than two rows of bony armor running its length, and by the form of the armor. They interpreted Rugosuchus as similar to Bernissartia , Shamosuchus and the unnamed "Glen Rose Form", but not as derived. [2] However, a more recent phylogenetic analysis by Turner and Buckley (2008) places it with the "Glen Rose Form" in a clade with Eusuchia, more derived than Bernissartia. [4]
Changchunsaurus is an extinct genus of small herbivorous dinosaur from Early Cretaceous deposits of Gongzhuling, Jilin, China. It is the first named dinosaur genus from Jilin.
Bernissartia is an extinct genus of neosuchian crocodyliform that lived in the Early Cretaceous, around 130 million years ago.
Chimaerasuchus is an extinct genus of Chinese crocodyliform from the Early Cretaceous Wulong Formation. The four teeth in the very tip of its short snout gave it a "bucktoothed" appearance. Due its multicusped teeth and marked heterodonty, it is believed to have been an herbivore. Chimaerasuchus was originally discovered in the 1960s but not identified as a crocodyliform until 1995, instead thought to possibly be a multituberculate mammal. It is highly unusual, as only two other crocodyliforms have displayed any characteristics resembling its adaptations to herbivory.
Mahajangasuchus is an extinct genus of crocodyliform which had blunt, conical teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar. It was a fairly large predator, measuring up to 4 metres (13 ft) long.
Pachycheilosuchus is an extinct genus of neosuchian from the Early Cretaceous of Texas, United States. Previously known, in part, as the "Glen Rose form", this crocodylomorph is notable for its procoelous vertebrae, otherwise found only in derived eusuchian crocodilians, a thick margin on the maxillae, and a shield of armor on the neck formed by the fusion of six individual scutes.
Junggarsuchus is an extinct genus of sphenosuchian crocodylomorph from the Middle or Late Jurassic period of China. The type and only species is J. sloani. The generic name of Junggarsuchus comes from the Junggar Basin, where the fossil was found, and the Greek word "souchos" meaning crocodile. The specific name, "sloani" is in honor of C. Sloan, who is credited with finding the holotype.
Gilchristosuchus is an extinct genus of neosuchian crocodyliform. Its fossils have been found in the upper Milk River Formation of Alberta, Canada, in rocks of either latest Santonian or earliest Campanian age. Gilchristosuchus was described in 1993 by Wu and Brinkman. The type species is G. palatinus, in reference to its distinctive palatine bones.
Iharkutosuchus is an extinct genus of basal eusuchian crocodyliform. Its fossils have been found in the Santonian-age Upper Cretaceous Csehbánya Formation in the Bakony Mountains of western Hungary.
Chiayusuchus is an extinct genus of stomatosuchid crocodylomorph. Fossils have been found from the Xinminbao Group in Gansu, China. It existed during the Barremian and Aptian stages of the Early Cretaceous. Although it was once considered as member of Stomatosuchidae, it is denied by having different teeth shape.
Isisfordia is an extinct genus of crocodyliform closely related to crocodilians that lived in Australia during the Middle Cretaceous (Albian–Cenomanian).
Kaprosuchus is an extinct genus of mahajangasuchid crocodyliform. It is known from a single nearly complete skull collected from the Upper Cretaceous Echkar Formation of Niger. The name means "boar crocodile" from the Greek κάπρος, kapros ("boar") and σοῦχος, soukhos ("crocodile") in reference to its unusually large caniniform teeth which resemble those of a boar. It has been nicknamed "BoarCroc" by Paul Sereno and Hans Larsson, who first described the genus in a monograph published in ZooKeys in 2009 along with other Saharan crocodyliformes such as Anatosuchus and Laganosuchus. The type species is K. saharicus.
Paralligator is an extinct genus of neosuchian crocodylomorph that lived during the Late Cretaceous (Cenomanian-Maastrichtian) period in what is now the Bayan Shireh and Nemegt formations of Mongolia, approximately 96 million to 70 million years ago. P. gradilifrons reached up to 4 m (13 ft) in length.
Shartegosuchidae is an extinct family of Late Jurassic and Early Cretaceous crocodyliforms. The family is named after the Late Jurassic Shar Teeg Beds in southwestern Mongolia, from which most shartegosuchid remains have been found. Five genera are currently assigned to Shartegosuchidae: Shartegosuchus, Nominosuchus, Kyasuchus, Adzhosuchus, and Fruitachampsa. Shartegosuchus, Nominosuchus, and Adzhosuchus all come from Shar Teeg, while Kyasuchus is known from the Early Cretaceous of Russia. Fruitachampsa is known from the Late Jurassic Morrison Formation of the western United States.
Shantungosuchus is an extinct genus of Early Cretaceous crocodyliform found in China. It includes three species: Shantungosuchus chuhsienensis and S. brachycephalus, which were both described by Yang Zhongjian – usually referred to as "Young" – in 1961 and 1982, and S. hangjinensis, which was described by Xiao-Chun Wu et al in 1994. S. chuhsienensis is the type for this genus.
Susisuchus is an extinct genus of neosuchian mesoeucrocodylian crocodyliform from the Early Cretaceous of Brazil. Fossils have been found from the Nova Olinda Member of the Aptian-age Crato Formation in the Araripe and Lima Campos Basins of northeastern Brazil. Named in 2003, Susisuchus is the sole member of the family Susisuchidae, and is closely related to the clade Eusuchia, which includes living crocodilians. The type species is S. anatoceps, known from a single partial articulated skeleton that preserves some soft tissue. A second species, S. jaguaribensis, was named in 2009 from fragmentary remains.
Pissarrachampsa is an extinct genus of baurusuchid mesoeucrocodylian from the Late Cretaceous of Brazil. It is based on a nearly complete skull and a referred partial skull and lower jaw from the ?Campanian - ?Maastrichtian-age Vale do Rio do Peixe Formation of the Bauru Group, found in the vicinity of Gurinhatã, Brazil.
Aegisuchus is an extinct monospecific genus of giant, flat-headed crocodyliform within the family Aegyptosuchidae. It was found in the Kem Kem Formation of southeast Morocco, which dates back to the Cenomanian age of the Late Cretaceous epoch. The type species Aegisuchus witmeri was named in 2012 by paleontologists Casey Holliday and Nicholas Gardner, who nicknamed it "Shieldcroc" for the shield-like shape of its skull. A. witmeri is known from a single partial skull including the braincase and skull roof.
Paluxysuchus is an extinct genus of neosuchian crocodyliform known from the Early Cretaceous Twin Mountains Formation of north-central Texas. It contains a single species, Paluxysuchus newmani. Paluxysuchus is one of three crocodyliforms known from the Early Cretaceous of Texas, the others being Pachycheilosuchus and an unnamed species referred to as the "Glen Rose Form". Paluxysuchus has a long, flat skull that is probably transitional between the long and narrow skulls of many early neosuchians and the short and flat skulls of later neosuchians.
Coelognathosuchia is an extinct clade of neosuchian crocodyliforms that includes all taxa more closely related to the family Pholidosauridae than to Bernissartia fagesii or Eusuchia. Martin et al. (2014) named the clade after finding goniopholidids and pholidosaurids to group together in their phylogenetic analysis of crocodyliform evolutionary relationships. In their analysis, Pholidosauridae was monophyletic and Goniopholididae was paraphyletic, being an assemblage of successively more basal taxa within Coelognathosuchia. Coelognathosuchia itself was positioned near the base of the larger clade Neosuchia as the sister group to a clade containing the Early Cretaceous neosuchian Bernissartia and Eusuchia, the group that includes all modern crocodilians and their closest extinct relatives.
Yanjisuchus is an extinct genus of paralligatorid neosuchian known from the Early Cretaceous Longjing Formation of Guizhou, China. It contains a single species, Y. longshanensis.