Allodaposuchidae | |
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A. precedens skull | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauria |
Clade: | Pseudosuchia |
Clade: | Crocodylomorpha |
Clade: | Crocodyliformes |
Clade: | Neosuchia |
Clade: | Eusuchia |
Clade: | † Allodaposuchidae Narváez et al., 2015 |
Type species | |
† Allodaposuchus precedens Nopcsa, 1928 | |
Genera | |
Allodaposuchidae is an extinct clade of eusuchians that lived in Europe during the Late Cretaceous (Santonian-Maastrichtian). [1] [2] [3]
The type genus, Allodaposuchus , was originally described in 1928 by Nopcsa from the Maastrichtian-age Sard Formation of the Hațeg Basin in Transylvania, Romania, and classified as a relative of the North American Leidyosuchus . [4] It was later classified as a eusuchian outside of Crocodylia in a 2001 paper, [5] and subsequent studies found a number of European eusuchian species ( Arenysuchus , Ischyrochampsa , Massaliasuchus , Musturzabalsuchus ) to group with Allodaposuchus, prompting the erection of the clade Allodaposuchidae to accommodate Allodaposuchus and all European eusuchians closely related to it. [2] [3] [6] [7] [8] Narváez et al. cladistically defined Allodaposuchidae in 2015 as Allodaposuchus precedens and all crocodyliforms more closely related to it than to Hylaeochampsa vectiana , Shamosuchus djadochtaensis , Borealosuchus sternbergii , Planocrania datangensis , Alligator mississippiensis , Crocodylus niloticus , or Gavialis gangeticus . [2] [9]
The exact placement of Allodaposuchidae is still in dispute. Narváez et al. considered it the sister group to Hylaeochampsidae, which together form a clade that is sister to Crocodylia. [2] Other studies have alternatively recovered them not as sister taxon, but rather as an evolutionary grade towards Crocodylia, with Hylaeochampsidae more basal than Allodaposuchidae. [10] [9] Alternatively, a 2021 analysis incorporating postcranial information recovered Allodaposuchidae within Crocodylia. [11]
Cladogram 1: Narváez et al., 2015
| Cladogram 2: Rio & Mannion, 2021
| Cladogram 3: Blanco, 2021
|
The internal phylogeny of Allodaposuchidae can be shown in the cladogram below from the 2021 Blanco study: [11]
Allodaposuchidae | |
In the 2021 study, Blanco recovered Allodaposuchus as paraphyletic, with Agaresuchus and Lohuecosuchus . Accordingly, Blanco proposed that Agaresuchus and Lohuecosuchus should be considered junior synonyms of Allodaposuchus. [11]
Allodaposuchus is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages, and possibly the Santonian stage, of the Late Cretaceous. Although generally classified as a non-crocodylian eusuchian crocodylomorph, it is sometimes placed as one of the earliest true crocodylians. Allodaposuchus is one of the most common Late Cretaceous crocodylomorphs from Europe, with fossils known from Romania, Spain, and France.
Boverisuchus is an extinct genus of planocraniid crocodyliforms known from the middle Eocene of Germany and western North America. It was a relatively small crocodyliform with an estimated total length of approximately 2.2–3.6 metres (7.2–11.8 ft).
Borealosuchus is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Christopher Brochu in 1997 for several species that had been assigned to Leidyosuchus. The species assigned to it are: B. sternbergii, the type species, from the Maastrichtian of Colorado, Montana, North Dakota, South Dakota, and Wyoming; B. acutidentatus, from the Paleocene of Saskatchewan; B. formidabilis, from the Paleocene of North Dakota; B. griffithi, from the Paleocene of Alberta; and B. wilsoni, from the Eocene of Wyoming. B. formidabilis is particularly well-known, represented by the remains of many individuals from the Wannagan Creek site in North Dakota. An indeterminate species is known from the Late Cretaceous Demopolis Chalk in Alabama.
Aegyptosuchus is an extinct monospecific genus of aegyptosuchid eusuchian crocodyliform. It was found in the Bahariya Formation of Egypt, which dates back to the Cenomanian age of the Late Cretaceous. The type and only species is Aegyptosuchus peyeri.
Eusuchia is a clade of neosuchian crocodylomorphs that first appeared in the Early Cretaceous, which includes modern crocodilians. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-Pg extinction.
Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more closely related to the alligators than the two other groups.
Acynodon is an extinct genus of eusuchian crocodylomorph from the Late Cretaceous, with fossils found throughout Southern Europe.
Massaliasuchus is an extinct monospecific genus of allodaposuchid eusuchian crocodyliform that is known from fossils found in Santonian–Campanian-age Upper Cretaceous rocks of southeastern France.
Brillanceausuchus is an extinct genus of paralligatorid crocodyliforms. Fossils have been found in Early Cretaceous–age rocks of Cameroon. The genus is notable for the position of the secondary choana within its palate. Parts of the pterygoid bones make up the rostral margin of the choana and thus separate it from the palatines, a feature also seen in the more advanced neosuchian suborder Eusuchia. This characteristic was once thought to be characteristic of Eusuchia, but its presence in Brillanceausuchus suggests that the trait is homoplasic, thus making the evolution of the position of the choana within crocodilians more complex than previously thought.
Dolichochampsa is an extinct genus of eusuchian crocodylomorph. It is the type genus and only member of the family Dolichochampsidae. Fossils have been found in the Yacoraite Formation of Argentina and the El Molino Formation of Bolivia of Maastrichtian age. It had a distinctive slender snout. Because the material associated with the specimens is so fragmentary, its relationships with other eusuchians remain unknown. Jouve et al. (2020) assigned Dolichochampsa to Gavialoidea, making it the oldest known South American member of this clade.
Musturzabalsuchus is an extinct monospecific genus of allodaposuchid eusuchian crocodyliform. The type and only species is Musturzabalsuchus buffetauti.
Planocrania is an extinct genus of eusuchian crocodyliforms from what is now China. Two species are currently known to belong to the genus.
Crocodyloidea is one of three superfamilies of crocodilians, the other two being Alligatoroidea and Gavialoidea, and it includes the crocodiles. Crocodyloidea may also include the extinct Mekosuchinae, native to Australasia from the Eocene to the Holocene, although this is disputed.
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial Gavialis gangeticus and the false gharial Tomistoma schlegelii are alive today, with Hanyusuchus having become extinct in the last few centuries.
Aegyptosuchidae is an extinct family of eusuchian crocodyliforms from the Cretaceous period of Africa. They are characterized by their large size and flat heads. The family includes two genera, Aegyptosuchus and Aegisuchus.
Planocraniidae is an extinct family of eusuchian crocodyliforms known from the Paleogene of Asia, Europe and North America. The family was coined by Li in 1976, and contains three genera, Boverisuchus, Duerosuchus and Planocrania. Planocraniids were highly specialized crocodyliforms that were adapted to living on land. They had extensive body armor, long legs, and blunt claws resembling hooves, and are sometimes informally called "hoofed crocodiles".
Lohuecosuchus is an extinct genus of allodaposuchid eusuchian crocodylomorph that lived during the Late Cretaceous in what is now Spain and southern France.
Agaresuchus is an extinct genus of allodaposuchid eusuchian crocodylomorph from the Late Cretaceous (Campanian-Maastrichtian) of Spain. It includes two species, the type species Agaresuchus fontisensis, and Agaresuchus subjuniperus, which was originally named as a species of the related genus Allodaposuchus. However, it has been proposed that both species may instead belong to the genus Allodaposuchus.
Longirostres is a clade of crocodilians that includes the crocodiles and the gavialids, to the exclusion of the alligatoroids. Defined in 2003 by Harshman et al., Longirostres is a crown group defined phylogenetically as including the last common ancestor of Crocodylus niloticus and Gavialis gangeticus and all of its descendants.
Aprosuchus is a genus of small-bodied Maastrichtian atoposaurid Eusuchian from the Hateg Basin, Romania.