Mandibular symphysis

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Mandibular symphysis
Symphysis menti (Gray190 edit).png
Anterior view of mandible, showing mandibular symphysis (red broken line)
Gray177.png
Medial surface of the left half of the mandible, dis-articulated from the right side at the mandibular symphysis
Details
Identifiers
Latin symphysis mandibulae
TA98 A02.1.15.004
TA2 838
FMA 75779
Anatomical terms of bone

In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: symphysis menti) or line of junction where the two lateral halves of the mandible typically fuse in the first year of life (6–9 months after birth). [1] It is not a true symphysis as there is no cartilage between the two sides of the mandible.

Contents

This ridge divides below and encloses a triangular eminence, the mental protuberance, the base of which is depressed in the center but raised on either side to form the mental tubercle. The lowest (most inferior) end of the mandibular symphysis — the point of the chin — is called the "menton". [2] [3]

It serves as the origin for the geniohyoid and the genioglossus muscles.

Other animals

Humpback skeleton showing the flexible "slingshot" symphysis present in baleen whales Humpback Whale skeleton.jpg
Humpback skeleton showing the flexible "slingshot" symphysis present in baleen whales
Skull and jaws of the "tetralophodont gomphothere" Tetralophodon longirostris, a proboscidean demonstrating mandibular symphysis elongation Tetralophodon longirostris skull - Batallones 2 fossil site, Torrejon de Velasco, Madrid, Spain.jpg
Skull and jaws of the "tetralophodont gomphothere" Tetralophodon longirostris, a proboscidean demonstrating mandibular symphysis elongation

Solitary mammalian carnivores that rely on a powerful canine bite to subdue their prey have a strong mandibular symphysis, while pack hunters delivering shallow bites have a weaker one. [4] When filter feeding, the baleen whales, of the suborder Mysticeti, can dynamically expand their oral cavity in order to accommodate enormous volumes of sea water. This is made possible thanks to its mandibular skull joints, especially the elastic mandibular symphysis which permits both dentaries to be rotated independently in two planes. This flexible jaw, which made the titanic body sizes of baleen whales possible, is not present in early whales and most likely evolved within Mysticeti. [5]

Many primitive proboscideans belonging to the group Elephantiformes have a greatly elongated mandibular symphysis. This was lost in many later groups, including modern elephants. [6]

Related Research Articles

<span class="mw-page-title-main">Baleen whale</span> Whales that strain food from the water using baleen

Baleen whales, also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea which use keratinaceous baleen plates in their mouths to sieve planktonic creatures from the water. Mysticeti comprises the families Balaenidae, Balaenopteridae (rorquals), Eschrichtiidae and Cetotheriidae. There are currently 16 species of baleen whales. While cetaceans were historically thought to have descended from mesonychians, molecular evidence instead supports them as a clade of even-toed ungulates (Artiodactyla). Baleen whales split from toothed whales (Odontoceti) around 34 million years ago.

<span class="mw-page-title-main">Neobalaenidae</span> Family of mammals

Neobalaenidae is a family of baleen whales including the extant pygmy right whale. Although traditionally considered related to balaenids, recent studies by Fordyce and Marx (2013) and Ludovic Dutoit and colleagues (2023) have recovered the living pygmy right whale as a member of Cetotheriidae, making it the only extant cetotheriid. Not all authors agree with this placement.

<span class="mw-page-title-main">Mental foramen</span> Opening on the anterior aspect of the mandible through which the mental nerve and vessels exit

The mental foramen is one of two foramina (openings) located on the anterior surface of the mandible. It is part of the mandibular canal. It transmits the terminal branches of the inferior alveolar nerve and the mental vessels.

<i>Thalassodromeus</i> Genus of azhdarchoid pterosaur from the Early Cretaceous

Thalassodromeus is a genus of pterosaur that lived in what is now Brazil during the Early Cretaceous period, about a hundred million years ago. The original skull, discovered in 1983 in the Araripe Basin of northeastern Brazil, was collected in several pieces. In 2002, the skull was made the holotype specimen of Thalassodromeus sethi by palaeontologists Alexander Kellner and Diogenes de Almeida Campos. The generic name means "sea runner", and the specific name refers to the Egyptian god Seth due to its crest being supposedly reminiscent of Seth's crown. Other scholars have pointed out that the crest was instead similar to the crown of Amon. A jaw tip was assigned to T. sethi in 2005, became the basis of the new genus Banguela in 2014, and assigned back to Thalassodromeus as the species T. oberlii in 2018. Another species was described in 2015 based on a supposed crest fragment, but this was later shown to be part of a turtle shell.

Peloneustes is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally described as a species of Plesiosaurus by palaeontologist Harry Govier Seeley in 1869, before being given its own genus by naturalist Richard Lydekker in 1889. While many species have been assigned to Peloneustes, P. philarchus is currently the only one still considered valid, with the others moved to different genera, considered nomina dubia, or synonymised with P. philarchus. Some of the material formerly assigned to P. evansi have since been reassigned to "Pliosaurus" andrewsi. Peloneustes is known from many specimens, including some very complete material.

<span class="mw-page-title-main">Chin</span> Facial feature

The chin is the forward pointed part of the anterior mandible below the lower lip. A fully developed human skull has a chin of between 0.7 cm and 1.1 cm.

<i>Janjucetus</i> Extinct genus of mammals

Janjucetus is an extinct genus of cetacean, and a basal baleen whale (Mysticeti), from the Late Oligocene around 25 million years ago (mya) off south-east Australia, containing one species J. hunderi. Unlike modern mysticetes, it possessed large teeth for gripping and shredding prey, and lacked baleen, and so was likely to have been a predator that captured large single prey animals rather than filter feeding. However, its teeth may have interlocked, much like those of the modern-day filter-feeding crabeater seal, which would have allowed some filter-feeding behaviour. Its hunting behaviour was probably similar to the modern-day leopard seal, probably eating large fish. Like baleen whales, Janjucetus could not echolocate; however, it did have unusually large eyes, and so probably had an acute sense of vision. The only specimen was found on the Jan Juc beach, where the remains of the extinct whales Mammalodon, Prosqualodon and Waipatia have also been discovered.

Mammalodon is an extinct genus of archaic baleen whale belonging to the family Mammalodontidae.

<span class="mw-page-title-main">Mammalodontidae</span> Extinct family of mammals

Mammalodontidae is a family of extinct whales known from the Oligocene of Australia and New Zealand.

<i>Aetiocetus</i> Extinct genus of mammals

Aetiocetus is a genus of extinct basal mysticete, or baleen whale that lived 33.9 to 23.03 million years ago, in the Oligocene in the North Pacific ocean, around Japan, Mexico, and Oregon, U.S. It was first described by Douglas Emlong in 1966 and currently contains known four species, A. cotylalveus, A. polydentatus, A. tomitai, and A. weltoni. These whales are remarkable for their retention of teeth and presence of nutrient foramina, indicating that they possessed baleen. Thus, Aetiocetus represents the transition from teeth to baleen in Oligocene mysticetes. Baleen is a highly derived character, or synapomorphy, of mysticetes, and is a keratinous structure that grows from the palate, or roof of the mouth, of the whale. The presence of baleen is inferred from the fossil record in the skull of Aetiocetus. Aetiocetus is known from both sides of the Pacific Ocean: it was first documented in Oregon, United States, but it is also known from Japan and Mexico. The genus is currently constrained to the Northern hemisphere and has little value in biostratigraphic studies of the Oligocene due to its limited occurrences across the Pacific.

The simian shelf is a bony thickening on the front of the ape mandible. Its function is to reinforce the jaw, though it also has the effect of considerably reducing the movement of the tongue by restricting the area available for muscles.

<i>Livyatan</i> Extinct genus of sperm whale from the Miocene epoch

Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.

<span class="mw-page-title-main">Cetotheriidae</span> Family of mammals

Cetotheriidae is a family of baleen whales. The family is known to have existed from the Late Oligocene to the Early Pleistocene before going extinct. Although some phylogenetic studies conducted by Fordyce & Marx 2013 recovered the living pygmy right whale as a member of Cetotheriidae, making the pygmy right whale the only living cetotheriid, other authors either dispute this placement or recover Neobalaenidae as a sister group to Cetotheriidae.

<span class="mw-page-title-main">Mandible</span> Lower jaw bone

In jawed vertebrates, the mandible, lower jaw, or jawbone is a bone that makes up the lower – and typically more mobile – component of the mouth.

<i>Llanocetus</i> Large Eocene whale from Antarctica and Bangladesh

Llanocetus is a genus of extinct toothed baleen whales from the Late Eocene of Antarctica. The type species, Llanocetus denticrenatus, reached gigantic proportions, with the juvenile specimen reaching an estimated 8 m (26 ft) in length; a second, unnamed species, known only from three isolated premolar teeth, reached an estimated total body length of up to 12 m (39 ft). Like other contemporary baleen whales of the Eocene, Llanocetus completely lacked baleen in its jaws. It was probably a suction feeder like the modern beaked and pygmy right whales.

Eomysticetidae is a family of extinct mysticetes belonging to Chaeomysticeti. It is one of two families in the basal chaeomysticete clade Eomysticetoidea.

<span class="mw-page-title-main">Elephantiformes</span> Suborder of mammals

Elephantiformes is a suborder within the order Proboscidea that contains the elephants. Members of this group are primitively characterised by the possession of an elongated mandibular symphysis and lower tusks. The earliest known member of the group, Dagbatitherium is known from the Eocene (Lutetian) of Togo, which is only known from isolated teeth, while other elephantiforms like Phiomia and Palaeomastodon are known from the Early Oligocene onwards. Most diversity of the group is placed in the subclade Elephantimorpha, which includes modern elephants, as well as other extinct groups like mastodons and gomphotheres.

<i>Waharoa <span style="font-style:normal;">(whale)</span></i> Extinct species of whale

Waharoa is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. It was identified with the discovery of Waharoa ruwhenua by Boessenecker and Fordyce (2015), which added a new genus and species to a monophyletic family Eomysticetidae.

<i>Coronodon</i> Extinct genus of whales

Coronodon is a genus of toothed (transitional) baleen whales from the Early Oligocene Ashley and Chandler Bridge formations of South Carolina. The genus contains three species: the type species C. havensteini, and additional species C. newtonorum and C. planifrons.

<i>Ankylorhiza</i> Extinct genus of toothed whales from the Oligocene epoch

Ankylorhiza is an extinct genus of toothed whale that lived in what is now the United States during the Oligocene epoch, between 29 and 23.5 million years ago. The type and only known species is A. tiedemani, though two fossil skeletons may represent an additional, second species within the genus. Ankylorhiza was about 4.8 meters (16 ft) long, with a long, robust skull bearing conical teeth that were angled forwards at the tip of the snout.

References

PD-icon.svgThis article incorporates text in the public domain from page 172 of the 20th edition of Gray's Anatomy (1918)

Notes

  1. Becker, Marshall Joseph (Apr 1986). "Mandibular symphysis (medial suture) closure in modern Homo sapiens: Preliminary evidence from archaeological populations". American Journal of Physical Anthropology. 69 (4): 499–501. doi:10.1002/ajpa.1330690409. ISSN   0002-9483 . Retrieved 2023-12-10.
  2. "Menton". The Free Dictionary. Retrieved 1 November 2016.
  3. Phulari, Basavaraj Subhashchandra (2013). An atlas on cephalometric landmarks (1st ed.). New Delhi: Jaypee Brothers Medical Publishers. p. 174. ISBN   9789350903247.
  4. Therrien, François (2005). "Mandibular force profiles of extant carnivorans and implications for the feeding behaviour of extinct predators". Journal of Zoology. 267 (3): 249. doi:10.1017/S0952836905007430.
  5. Fitzgerald 2012
  6. Li, Chunxiao; Deng, Tao; Wang, Yang; Sun, Fajun; Wolff, Burt; Jiangzuo, Qigao; Ma, Jiao; Xing, Luda; Fu, Jiao (2023-08-16). Longer mandible or nose? Co-evolution of feeding organs in early elephantiforms (Report). Paleontology. doi:10.1101/2023.08.15.553347.

Sources