Sandpiper

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Sandpipers
Temporal range: Early Oligocene to recent
Catoptrophorus semipalmatus edit.jpg
Willet (Tringa semipalmata)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Charadriiformes
Suborder: Scolopaci
Family: Scolopacidae
Rafinesque, 1815
Type genus
Scolopax
Genera

Scolopacidae is a large family of shorebirds, or waders, which mainly includes many species known as sandpipers, but also others such as woodcocks, curlews and snipes. The majority of these species eat small invertebrates picked out of the mud or soil. Different lengths of bills enable different species to feed in the same habitat, particularly on the coast, without direct competition for food.

Contents

Sandpipers have long bodies and legs, and narrow wings. Most species have a narrow bill, but otherwise the form and length are quite variable. They are small to medium-sized birds, measuring 12 to 66 cm (4.7–26.0 in) in length. The bills are sensitive, allowing the birds to feel the mud and sand as they probe for food. They generally have dull plumage, with cryptic brown, grey, or streaked patterns, although some display brighter colours during the breeding season. [1]

Most species nest in open areas, and defend their territories with aerial displays. The nest itself is a simple scrape in the ground, in which the bird typically lays three or four eggs. The young of most species are precocial. [1]

Sandpiper nest with four eggs Sandpiper nest with four eggs.jpg
Sandpiper nest with four eggs

Taxonomy

The family Scolopacidae was introduced (as Scolopacea) by the French polymath Constantine Samuel Rafinesque in 1815. [2] [3] The family contains 98 extant or recently extinct species divided into 15 genera. [4] For more details, see the article List of sandpiper species.

The following genus level cladogram of the Scolopacidae is based on a study by David Černý and Rossy Natale that was published in 2022. [5]

Scolopacidae

Bartramia – upland sandpiper

Numenius – curlews (9 species)

Limosa – godwits (4 species)

Limnodromus – dowitchers (3 species)

Lymnocryptes – jack snipe

Scolopax – woodcocks (8 species)

Gallinago imperialis – imperial snipe

Gallinago undulata – giant snipe

Coenocorypha – austral snipes (3 extant and 6 extinct species)

Gallinago – snipes (18 species)

Xenus – Terek sandpiper

Actitis – sandpipers (2 species)

Phalaropus – phalaropes (3 species)

Tringa – sandpipers, shanks, tattlers etc (13 species)

Prosobonia – Polynesian sandpipers (1 extant and 3 extinct species)

Arenaria – turnstones (2 species)

Calidris – sandpipers (24 species)


ImageGenusLiving species
CURLEW, LONG-BILLED (3-22-10) morro bay, ca -04 (4455146301).jpg Numenius Brisson, 1760
Bar-tailed Godwit.jpg Limosa Brisson, 1760
Arenaria interpres (habitus).jpg Arenaria Brisson, 1760
Tuamotu sandpiper.jpg Prosobonia Bonaparte, 1850
Semipalmated Sandpiper (8578570180).jpg Calidris Merrem, 1804
Short-billed dowitcher in JBWR (40844).jpg Limnodromus Wied-Neuwied, 1833
EURASIAN-WOODCOCK-MANGPOO.jpg Scolopax Linnaeus, 1758
Coenocorypha aucklandica meinertzhagenae 387315409 (cropped).jpg Coenocorypha G.R. Gray, 1855
Jack snipe.png Lymnocryptes F. Boie, 1826
Gallinago gallinago a1.JPG Gallinago Brisson, 1760
Terek sandpiper 9.jpg Xenus Kaup, 1829
  • Terek sandpiper (Xenus cinereus)
Phalaropus fulicarius 98755138 (cropped).jpg Phalaropus Brisson, 1760
Actitis hypoleucos 1 tb (Marek Szczepanek).jpg Actitis Illiger, 1811
Lesser yellowlegs bunche beach (31791842132).jpg Tringa Linnaeus, 1758

Evolution

The early fossil record is scant for a group that was probably present at the non-avian dinosaur's extinction. "Totanus" teruelensis (Late Miocene of Los Mansuetos (Spain) is sometimes considered a scolopacid – maybe a shank – but may well be a larid; little is known of it.

Paractitis has been named from the Early Oligocene of Saskatchewan (Canada), while Mirolia is known from the Middle Miocene at Deiningen in the Nördlinger Ries (Germany). Most living genera would seem to have evolved throughout the Oligocene to Miocene with the waders perhaps a bit later; see the genus accounts for the fossil record.

In addition there are some indeterminable remains that might belong to extant genera or their extinct relatives:

Description

The least sandpiper is the smallest species of sandpiper Least Sandpiper Don Edwards WR 1.jpg
The least sandpiper is the smallest species of sandpiper

The sandpipers exhibit considerable range in size and appearance, the wide range of body forms reflecting a wide range of ecological niches. Sandpipers range in size from the least sandpiper, at as little as 18 grams (0.040 pounds) and 11 cm (4.3 in) in length, to the Far Eastern curlew, at up to 66 cm (26 in) in length, and the Eurasian curlew, at up to 1.3 kg (2.9 lb). Within species there is considerable variation in patterns of sexual dimorphism. Males are larger than females in ruffs and several sandpipers, but are smaller than females in the knots, curlews, phalaropes and godwits. The sexes are similarly sized in the snipes, woodcock and tringine sandpipers. Compared to the other large family of wading birds, the plovers (Charadriidae), they tend to have smaller eyes, more slender heads, and longer thinner bills. Some are quite long-legged, and most species have three forward pointing toes with a smaller hind toe (the exception is the sanderling, which lacks a hind toe). [9]

Sandpipers are more geared towards tactile foraging methods than the plovers, which favour more visual foraging methods, and this is reflected in the high density of tactile receptors in the tips of their bills. These receptors are housed in a slight horny swelling at the tip of the bill (except for the surfbird and the two turnstones). Bill shape is highly variable within the family, reflecting differences in feeding ecology. Bill length relative to head length varies from three times the length of the head in the long-billed curlew to just under half the head length in the Tuamotu sandpiper. Bills may be straight, slightly upcurled or strongly downcurved. [9] Like all birds, the bills of sandpipers are capable of cranial kinesis, literally being able to move the bones of the skull (other than the obvious movement of the lower jaw) and specifically bending the upper jaw without opening the entire jaw, an act known as rhynchokinesis. It has been hypothesized this helps when probing by allowing the bill to be partly opened with less force and improving manipulation of prey items in the substrate. Rhynchokinesis is also used by sandpipers feeding on prey in water to catch and manipulate prey. [10]

Distribution, habitat, and movements

Sandpipers spending the non-breeding season in Roebuck Bay, Western Australia Waders in flight Roebuck Bay.jpg
Sandpipers spending the non-breeding season in Roebuck Bay, Western Australia

The sandpipers have a cosmopolitan distribution, occurring across most of the world's land surfaces except for Antarctica and the driest deserts. A majority of the family breed at moderate to high latitudes in the Northern Hemisphere, in fact accounting for the most northerly breeding birds in the world. Only a few species breed in tropical regions, ten of which are snipes and woodcocks and the remaining species being the unusual Tuamotu sandpiper, which breeds in French Polynesia (although prior to the arrival of humans in the Pacific there were several other closely related species of Polynesian sandpiper). [9]

Diet and feeding

There are broadly four feeding styles employed by the sandpipers, although many species are flexible and may use more than one style. The first is pecking with occasional probing, usually done by species in drier habitats that do not have soft soils or mud. The second, and most frequent, method employed is probing soft soils, muds and sands for prey. The third, used by Tringa shanks, involves running in shallow water with the bill under the water chasing fish, a method that uses sight as well as tactile senses. The final method, employed by the phalaropes and some Calidris sandpipers, involves pecking at the water for small prey. [9] A few species of scolopacids are omnivorous to some extent, taking seeds and shoots as well as invertebrates.

Breeding

Many sandpipers form monogamous pairs, but some sandpipers have female-only parental care, some male-only parental care, some sequential polyandry and other compete for the mate on the lek. Sandpipers lay three or four eggs into the nest, which is usually a vague depression or scrape in the open ground, scarcely lined with soft vegetation. [9] In species where both parents incubate the eggs, females and males share their incubation duties in various ways both within and between species. In some pairs, parents exchange on the nest in the morning and in the evening so that their incubation rhythm follows a 24-hour day, in others each sex may sit on the nest continuously for up to 24 hours before it is exchanged by its partner. [11] In species where only a single parent incubates the eggs, during the night the parent sits on the eggs nearly continuously and then during the warmest part of a day leaves the nest for short feeding bouts. [12] Chicks hatch after about three weeks of incubation and are able to walk and forage within a few hours of hatching. A single parent or both parents guide and brood the chicks. [9]

Footnotes

  1. ↑ A distal right tarsometatarsus of a bird roughly similar to a pectoral sandpiper. Probably calidrid or basal to them, somewhat reminiscent of turnstones. [8]

Related Research Articles

<span class="mw-page-title-main">Phalarope</span> Genus of birds

A phalarope is any of three living species of slender-necked shorebirds in the genus Phalaropus of the bird family Scolopacidae.

<span class="mw-page-title-main">Turnstone</span> Genus of birds

Turnstones are two bird species that constitute the genus Arenaria in the family Scolopacidae. They are closely related to calidrid sandpipers and might be considered members of the tribe Calidriini.

<span class="mw-page-title-main">Ruddy turnstone</span> Species of bird

The ruddy turnstone is a small cosmopolitan wading bird, one of two species of turnstone in the genus Arenaria.

<i>Tringa</i> Genus of birds

Tringa is a genus of waders, containing the shanks and tattlers. The genus name Tringa is the Neo-Latin name given to the green sandpiper by the Italian naturalist Ulisse Aldrovandi in 1599. They are mainly freshwater birds, often with brightly coloured legs as reflected in the English names of six species, as well as the specific names of two of these and the green sandpiper. They are typically associated with northern hemisphere temperate regions for breeding. Some of this group—notably the green sandpiper—nest in trees, using the old nests of other birds, usually thrushes.

<span class="mw-page-title-main">Long-billed dowitcher</span> Species of bird

The long-billed dowitcher is a medium-sized shorebird with a relatively long bill belonging to the sandpiper family, Scolopacidae. In breeding plumage, adults are characterized by a beautiful rufous head and underparts with a darker mottled back and a large white upper rump only seen in flight. They feed in various freshwater habitats with their bill underwater in a "sewing machine" motion and are known to have an exciting mating display where males chase females in flight. The genus, Limnodromus is Ancient Greek from limne, "marsh" and dromos, "racer". The specific scolopaceus is Neo-Latin for "snipe-like", from Latin scolopax, scolopacis, a snipe or woodcock. The English name is from Iroquois and was first recorded in 1841.

<span class="mw-page-title-main">Long-billed curlew</span> Species of bird

The long-billed curlew is a large North American shorebird of the family Scolopacidae. This species was also called "sicklebird" and the "candlestick bird". The species breeds in central and western North America, migrating southward and coastward for the winter.

<span class="mw-page-title-main">Eskimo curlew</span> Species of bird

The Eskimo curlew, also known as northern curlew, is a species of curlew in the family Scolopacidae. It was one of the most numerous shorebirds in the tundra of western Arctic Canada and Alaska. Thousands of birds were then killed per year in the late 1800s. As there has not been a reliable sighting since 1987 or a confirmed sighting since 1963, the Eskimo curlew is considered Critically Endangered or possibly extinct. The bird is about 30 cm (12 in) long and fed mostly on insects and berries.

<span class="mw-page-title-main">Pin-tailed snipe</span> Species of bird

The pin-tailed snipe or pintail snipe is a species of bird in the family Scolopacidae, the sandpipers.

<span class="mw-page-title-main">Austral snipe</span> Genus of birds

The austral snipes, also known as the New Zealand snipes or tutukiwi, are a genus, Coenocorypha, of tiny birds in the sandpiper family, which are now only found on New Zealand's outlying islands. There are currently three living species and six known extinct species, with the Subantarctic snipe having three subspecies, including the Campbell Island snipe discovered as recently as 1997. The genus was once distributed from Fiji, New Caledonia and Norfolk Island, across New Zealand and southwards into New Zealand's subantarctic islands, but predation by introduced species, especially rats, has drastically reduced their range.

References

  1. 1 2 Harrison, Colin J.O. (1991). Forshaw, Joseph (ed.). Encyclopaedia of Animals: Birds. London: Merehurst Press. pp. 103–105. ISBN   1-85391-186-0.
  2. ↑ Rafinesque, Constantine Samuel (1815). Analyse de la nature ou, Tableau de l'univers et des corps organisés (in French). Vol. 1815. Palermo: Self-published. p. 70.
  3. ↑ Bock, Walter J. (1994). History and Nomenclature of Avian Family-Group Names. Bulletin of the American Museum of Natural History. Vol. 222. New York: American Museum of Natural History. pp. 113, 252. hdl:2246/830.
  4. ↑ Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2021). "Sandpipers, snipes, coursers". IOC World Bird List Version 11.2. International Ornithologists' Union. Retrieved 18 November 2021.
  5. ↑ Černý, David; Natale, Rossy (2022). "Comprehensive taxon sampling and vetted fossils help clarify the time tree of shorebirds (Aves, Charadriiformes)" (PDF). Molecular Phylogenetics and Evolution. 177: 107620. doi:10.1016/j.ympev.2022.107620. PMID   36038056.
  6. ↑ "Magornitho".
  7. ↑ "great Alaska department of fish and game".
  8. ↑ Wetmore, Alexander (1937). "The Eared Grebe and other Birds from the Pliocene of Kansas" (PDF). Condor . 39 (1): 40. doi:10.2307/1363487. JSTOR   1363487.
  9. 1 2 3 4 5 6 Piersma, Theunis (1996). "Family Scolopacidae (Snipes, Sandpipers and Phalaropes)" . In del Hoyo, Josep; Elliott, Andrew; Sargatal, Jordi (eds.). Handbook of the Birds of the World. Volume 3, Hoatzin to Auks. Barcelona: Lynx Edicions. pp. 444–487. ISBN   978-84-87334-20-7.
  10. ↑ Estrella, Sora; Masero, José A. (2007). "The use of distal rhynchokinesis by birds feeding in water". Journal of Experimental Biology. 210 (21): 3757–3762. doi: 10.1242/jeb.007690 . PMID   17951416.
  11. ↑ Bulla, Martin; Valcu, Mihai; Dokter, Adriaan M.; Dondua, Alexei G.; Kosztolányi, András; Rutten, Anne L.; Helm, Barbara; Sandercock, Brett K.; Casler, Bruce (2016). "Unexpected diversity in socially synchronized rhythms of shorebirds" (PDF). Nature. 540 (7631): 109–113. Bibcode:2016Natur.540..109B. doi:10.1038/nature20563. PMID   27880762. S2CID   4390453.
  12. ↑ Løfaldli, Lars (1985-01-01). "Incubation Rhythm in the Great Snipe Gallinago media". Holarctic Ecology. 8 (2): 107–112. Bibcode:1985Ecogr...8..107L. doi:10.1111/j.1600-0587.1985.tb01160.x. JSTOR   3682650.
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