Pterotracheoidea | |
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A pterotracheid | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Gastropoda |
Subclass: | Caenogastropoda |
Order: | Littorinimorpha |
Superfamily: | Pterotracheoidea Rafinesque, 1814 |
Families | |
See text | |
Synonyms | |
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The Pterotracheoidea is, according to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), a taxonomic superfamily of sea snails or sea slugs, marine gastropod molluscs in the clade Littorinimorpha. They are commonly called heteropods or sea elephants.
According to the World Registry of Marine Species, this superfamily comprises five families, two of which are extinct: [1]
These holoplanktonic snails are found floating or swimming in tropical to subtropical open oceans and seas at a depth of maximum 200 to 300 m [2]
These snails have adapted themselves to a pelagic living :
The taenioglossate radula, situated at the tip of the proboscis, can be protruded to capture the prey.
They have paired, large spherical eyes, contained in a gelatinous mass, that they use to locate their prey. As the body is transparent, one can easily see the internal organs and the statocysts with its statoliths (an organ that tells the animal whether it is upside down or not). The swimming fin of the foot has a ventral or posteroventral sucker. This sucker has grown larger in the family Atlantidae where it serves to hold the prey. The fin sucker is only present in male snails of the Pterotracheidae.
The body size varies from microscopic (Atlantidae) to macroscopic (Carinariidae and Pterotracheidae). Fertilized eggs are laid in mucous strings that hatch after a few days into free swimming veliger larvae (except in Firoloida desmaresti where the eggs remain attached in a tubular filament to the female body).
All the heteropods float or swim with their ventral part upward. The atlantids are negatively buoyant, while the others have neutral buoyancy.
These snails all have a coiled shell present in their larval stage. But the shell is no longer present in the Pterotracheidae after metamorphosis. The keel (sharp ridge on a whorl of the shell) of the adult shell extends outwards in the Atlantidae and anteriorly in Carinariidae. The shell is calcareous in the Carinariidae. In the Atlantidae, however, the shell and the keel can be calcareous (genus Atlanta) or the shell calcareous and its keel composed of conchiolin (genus Protatlanta), or the shell composed exclusively of conchiolin (genus Oxygyrus). Only the Atlantidae can retract into their shell.
The Thecosomata, or sea butterflies, are a taxonomic suborder of small, pelagic, free-swimming sea snails known as holoplanktonic opisthobranch gastropod mollusks, in the order Pteropoda. Most pteropods have some form of calcified shell, although it is often very light, even translucent.
Sea angels are a large group of small free-swimming sea slugs, classified into six separate families. They are pelagic opisthobranchs in the clade Gymnosomata within the larger mollusc clade Heterobranchia. Sea angels were previously considered to be pteropods.
Limacina is a genus of swimming predatory sea snails commonly known as sea butterflies in the family Limacinidae. This genus contains some of the world's most abundant gastropod species.
The superfamily Cavolinioidea is the most speciose group of sea butterflies. They belong to the suborder Euthecosomata.
Littorinimorpha is a large order of snails, gastropods, consisting primarily of sea snails, but also including some freshwater snails and land snails.
Ceolodiscus is a genus in the gastropod family Euomphalidae from the Lower and Middle Jurassic of Europe, mostly on Germany, France and United Kingdom. Other posible records include specimens from Hungary of Earliest Jurassic (Hettangian) age. As well there are specimens from Switzerland of Middle Jurassic age. It is the only genus in the monotypic family Coelodiscidae. The genus is usually allied with modern Pterotracheoidea, based mostly on it´s resemblance with modern Atlanta larvae, yet it differs by lacking extant family affiliations. This genus is linked with the Toarcian Oceanic Anoxic Event, that likely triggered it´s evolution.
Atlantidae is a family of sea snails, holoplanktonic gastropod molluscs in the clade Littorinimorpha.
Carinariidae, known by the common name "heteropods" like their relatives in the Pterotracheoidea, is a taxonomic family of swimming or floating sea snails, pelagic marine gastropod molluscs in the clade Littorinimorpha.
Carinaria is a genus of medium-sized floating sea snails, pelagic gastropod molluscs in the family Carinariidae.
Pterotracheidae is a family of medium-sized to large floating sea snails, pelagic gastropod molluscs. They are in the superfamily Pterotracheoidea along with two other similar pelagic families, the Atlantidae and the Carinariidae.
Vetigastropoda is a major taxonomic group of sea snails, marine gastropod mollusks that form a very ancient lineage. Taxonomically the Vetigastropoda are sometimes treated as an order, although they are treated as an unranked clade in Bouchet and Rocroi, 2005.
Cymbulioidea is a taxonomic superfamily of pelagic "sea butterflies", one group of swimming sea snails. They are holoplanktonic opisthobranch gastropod molluscs in the clade Thecosomata.
Atlanta echinogyra is a species of sea snail, a holoplanktonic marine gastropod mollusk in the family Atlantidae.
Oxygyrus keraudrenii is a species of sea snail, a holoplanktonic marine gastropod mollusk in the family Atlantidae.
Protatlanta souleyeti is a species of sea snail, a holoplanktonic marine gastropod mollusk in the family Atlantidae.
Atlanta is a genus of pelagic marine gastropod molluscs in the family Atlantidae.
Protatlanta rotundata is an extinct species of sea snail, a holoplanktonic marine gastropod mollusk in the family Atlantidae.
Limacina retroversa is a distinct species of swimming planktonic gastropods, belonging to a group of predatory sea snails known as sea butterflies (Thecosomata). The name Limacina retroversa describes the unique morphology of this sea snail, including its slug-like body and coiled, backwards-turning shell. They are typically found in the epipelagic zone of cold, polar waters, but can be found worldwide, in any ocean. L. retroversa are currently under threat, as their numbers are decreasing due to rising global carbon levels and other human-caused climate threats.
Carinaria cristata, commonly known as the glassy nautilus, is a species of pelagic marine gastropod mollusc in the family Carinariidae. It is found in the Pacific Ocean and is described as being holoplanktonic, because it spends its entire life as part of the plankton. It was first described by Carl Linnaeus in 1767. Its fragile shell was much prized by early conchologists for their collections, being so rare that it was said to be worth more than its weight in gold.
Carinaria galea, common name the helmeted carinaria, is a species of sea snail, a marine pelagic marine gastropod mollusc in the family Carinariidae. It was first described in 1835 by William Henry Benson, an amateur malacologist in the Bengal Civil Service.