Agnostida is an order of arthropod which first developed near the end of the Early Cambrian period and thrived during the Middle Cambrian. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders. The last agnostids went extinct in the Late Ordovician.
Trilobites are a group of extinct marine artiopodan arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period, and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 300 million years.
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
The thorax or chest is a part of the anatomy of humans, mammals, other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the creature's body, each of which is in turn composed of multiple segments.
The pygidium is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.
Harpetida is one of the nine orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period.
Phacopida ("lens-face") is an order of trilobite that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse group of related suborders.
Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.
Corynexochida is an order of trilobite that lived from the Lower Cambrian to the Late Devonian. Like many of the other trilobite orders, Corynexochida contains many species with widespread characteristics.
Proetida is an order of trilobite that lived from the Ordovician to the Permian. It was the last order of trilobite to go extinct, finally dying out in the Permian extinction.
Olenellina is a suborder of the order Redlichiida of Trilobites that occurs about halfway during the Lower Cambrian, at the start of the stage called the Atdabanian. The earliest trilobites in the fossil record are arguably Olenellina, although the earliest Redlichiina and Eodiscina follow quickly. The suborder died out when the Lower passed into the Middle Cambrian, at the end of the stage called Toyonian. A feature uniting the Olenellina is the lack of rupture lines in the headshield, which in other trilobites assist the periodic moulting, associated with arthropod growth. Some derived trilobites have lost facial sutures again, but all of these are blind, while all Olenellina have eyes.
Redlichiina is a suborder of the order Redlichiida of Trilobites. The suborder contains three superfamilies: Emuelloidea, Redlichioidea and Paradoxidoidea. These trilobites are some of the oldest trilobites known. They originated at the beginning of the Cambrian Period and disappeared at the end of the middle Cambrian.
Elliptocephala is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived from the later part of the Tommotian to the upper Botomian. Elliptocephala can easily be confused with Ellipsocephalus, a trilobite genus of the Ptychopariida order.
Aulacopleura is a genus of proetid trilobite that lived from the Middle Ordovician to the Middle Devonian. Some authors may classify this group as subgenus Otarion (Aulacopleura). The cephalon is semicircular or semielliptical, with border and preglabellar field. The glabella is short, with or without defined eye ridges connecting it with eyes of variable size. Spines at the rear outer corners of the cephalon are present, typically reaching back to the 2nd to 4th thorax segment. The 'palate' is not connected to the dorsal shield of the cephalon. The cephalon is pitted, or has small tubercles. The thorax has up to 22 segments. The pleural ends are usually rounded. The pygidium is small (micropygous), with an even margin. A. koninckii had a modern type of compound eye.
Ellipsocephalus is an extinct genus of blind trilobite that lived during the Cambrian. The genus comprises benthic species inhabiting deep, poorly lit or aphotic habitats. E. hoffi is a common trilobite mainly from central Europe.
Ptychopariidae is a family of trilobites, containing the following genera:
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.
Cyclopygidae is a family of asaphid trilobites from the Ordovician. Cyclopygids had an extratropical distribution, and there is evidence that they lived in darker parts of the water column. Cyclopygids are characterized by enlarged eyes, with a wide angle of view, both horizontal and vertical, reminiscent of the eyes of dragonflies. These typically touch the glabella directly on the side. Cyclopygids all lack genal spines, but Symphysops carries a forward directed frontal spine on the glabella. It is presumed that at least the members of the genus Pricyclopyge swam upside down and had bioluminescent organs on the third thorax segment. Cyclopygids had between 7 and 5 thorax segments, a wide and stout axis, and short side lobes.
Olenina is an extinct suborder of the trilobite order Ptychopariida.
Ptychoparioidea is a superfamily of the Ptychopariida order of trilobites.
