| Taldycupedidae Temporal range: | |
|---|---|
 | |
| Taldycupes reticulatus elytra photograph | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Coleoptera |
| Family: | † Taldycupedidae Rohdendorf, 1961 |
| Genera | |
See text | |
Taldycupedidae is an extinct family of beetles primarily known from the Permian period.
Kirejtshuk (2020) included the following genera in Taldycupedidae, preliminarily treating it as a subfamily Taldycupedinae of the family Permocupedidae. [1]
The genera Alveolacupes Lin, 1983, Clathropenna Fujiyama, 1973 and Penecupes Ren, 1995 were previously placed in the family by some authors, but were all excluded from it by Kirejtshuk (2020). The genera Wuchangia Hong, 1985 and Yuxianocoleus Hong, 1985, both from the Lower Jurassic of Hubei, China, were also excluded from the family by Zhao et al. (2021) (as the subfamily Taldycupedinae). [2]
The Permian is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia.
The Trachypachidae are a family of beetles that generally resemble small ground beetles, but that are distinguished by the large coxae of their rearmost legs. There are only six known extant species in the family, with four species of Trachypachus found in northern Eurasia and northern North America, and two species of Systolosoma in Chile and Argentina. They were much more diverse in the past, with dozens of described species from the Mesozoic.
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian.
The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic.
In the geologic timescale, the Capitanian is an age or stage of the Permian. It is also the uppermost or latest of three subdivisions of the Guadalupian Epoch or Series. The Capitanian lasted between 264.28 and 259.51 million years ago. It was preceded by the Wordian and followed by the Wuchiapingian.
In the geologic timescale, the Roadian is an age or stage of the Permian. It is the earliest or lower of three subdivisions of the Guadalupian Epoch or Series. The Roadian lasted between 273.01 and 266.9 million years ago (Ma). It was preceded by the Kungurian and followed by the Wordian.
In the geologic timescale, the Wuchiapingian or Wujiapingian is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between 259.51 and 254.14 million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian.
In the geologic time scale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian Epoch or Series. The Changhsingian lasted from 254.14 to 251.9 Ma ago. It is preceded by the Wuchiapingian age/stage and is followed by the Induan age/stage.
Kunpania is an extinct genus of dicynodont therapsid from the Quanzijie Formation of Xinjiang, China. The type and only species is K. scopulusa, and it is known only by a single incomplete specimen including parts of the skull and forelimb. Since its initial description in 1978 by palaeontologist Ailing Sun, it has sometimes been considered to be another species of Dicynodon by other researchers, or potentially undiagnostic. However, a redescription in 2021 reaffirmed its distinctiveness, including a uniquely well developed muscle attachment on the humerus. Kunpania is perhaps the oldest known member of the derived dicynodont group Dicynodontoidea, potentially dating to the Middle Permian period during the Capitanian, and so may fill a knowledge gap in the history of dicynodont evolution.
The Protocoleoptera are a paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern beetle groups during the following Triassic. Protocoleopterans typically possess prognathous (horizontal) heads, distinctive elytra with regular window punctures, culticles with tubercles or scales, as well as a primitive pattern of ventral sclerites, similar to the modern archostematan families Ommatidae and Cupedidae. They are thought to have been xylophagous and wood boring.
Permosynidae is an extinct family of beetles known from the Permian to Cretaceous. It is sometimes treated as a form family, because all known members were described from fossils of isolated elytra with punctate striae.
Platycrossos is an extinct genus of protocoleopteran beetles in the family Permosynidae. It is known from the Triassic and Jurassic of Australia, Austria, Mongolia and Russia. Like other members of the family, the species of this genus were described from fossils of isolated elytra with punctate striae.
Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one species known from the Triassic, Frankencupes ultimus from the Middle Triassic (Anisian) Röt Formation of Germany. They are thought to have been xylophagous, which is presumed to be the ancestral ecology of beetles.
Conodonts are an extinct class of animals whose feeding apparatuses called teeth or elements are common microfossils found in strata dating from the Stage 10 of the Furongian, the fourth and final series of the Cambrian, to the Rhaetian stage of the Late Triassic. These elements can be used alternatively to or in correlation with other types of fossils in the subfield of the stratigraphy named biostratigraphy.
Ademosynidae is an extinct family of beetles, known from the Late Triassic to Early Cretaceous. There are at least six genera and 11 species in Ademosynidae. Members of the family were small oval beetles, with a length generally smaller than 1 cm. Characteristics of the family include a pronotum without anterior angles and a rounded anterior margin, and elytra with 9–12 punctate striae. The systematic position of the family within the order Coleoptera is currently uncertain: they can be considered early examples of the suborder Polyphaga, or unusual examples of Archostemata.
Mesopsychidae is an extinct family of scorpionflies known from the Late Permian to Mid Cretaceous. It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Lophioneurida is an extinct order of Thysanoptera, dating from the Carboniferous to the Cretaceous. It is likely paraphyletic, with modern thrips having evolved from members of the group.
Tshekardocoleidae is an extinct family of beetles, known from the Permian. They represent amongst the oldest beetles. Like other primitive beetles, they are thought to have been xylophagous. They first appeared during the Cisuralian, before becoming extinct at the beginning of the Guadalupian. A claimed Jurassic record is doubtful. The oldest known beetle, Coleopsis, was originally assigned to this family, but is now assigned to its own family Coleopsidae.
Geinitziidae is an extinct family of polyneopteran insects, known from the Permian to Cretaceous. They are currently considered to be members "Grylloblattida" a poorly defined group of extinct insects thought to be related to modern ice crawlers (Grylloblattidae). Other authors place them in the extinct order Reculida. Unlike modern ice crawlers, which are wingless, they had large wings, bearing a superficial resemblance to cockroaches, and are thought to have been day-active above ground predators.
The Khuff Formation is a geologic formation in Oman and Saudi Arabia. It preserves fossils dating back to the Permian and early Triassic periods.