Taldycupedidae

Taldycupedidae Temporal range: Middle Permian–Late Permian PreꞒ Ꞓ O S D C P T J K Pg N
Taldycupes reticulatus elytra photograph
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	† Protocoleoptera
Superfamily:	† Permocupedoidea
Family:	† Taldycupedidae Rohdendorf, 1961
Genera
See text

Taldycupedidae is an extinct family of beetles primarily known from the Permian period.

Taxonomy

Kirejtshuk (2020) included the following genera in Taldycupedidae, preliminarily treating it as a subfamily Taldycupedinae of the family Permocupedidae. ^[1]

• Afrotaldycupes Kirejtshuk, 2020 – Beaufort Group, South Africa, Late Permian (Changhsingian)
  • Afrotaldycupes africanus(Ponomarenko, 2005) (Type species)
  • Afrotaldycupes lidgettoniensis(Ponomarenko, 2005)
• Mesothoris Tillyard, 1916 – Blackstone Formation, Australia Late Triassic (Norian)
  • Mesothoris clathrataTillyard, 1916 (Type species)
  • Mesothoris grandisDunstan, 1923
  • Mesothoris punctomarginumDunstan, 1923
• Proterocupes Ponomarenko, 2015 (syn LobanoviaKirejtshuk, 2020)
  • Proterocupes majorPonomarenko, 2015 – Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
  • Proterocupes nedubrovensisPonomarenko, 2015 – Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
  • Proterocupes permiana(Ponomarenko, 2015) (Type species) – Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
• Simmondsia Dunstan, 1923 – Blackstone Formation, Australia Late Triassic (Norian)
  • Simmondsia cylindricaDunstan, 1923
  • Simmondsia subpiriformisDunstan, 1923 (Type species)
• Taldycupes Rohdendorf, 1961 (syn CryptocupesRohdendorf, 1961, TaldycupidiumRohdendorf, 1961)
  • Taldycupes bergi(Rohdendorf, 1961) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Taldycupes cellulosusPonomarenko, 2013 – Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
  • Taldycupes khalfiniRohdendorf, 1961 (Type species) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Taldycupes moltshanoviRohdendorf, 1961 – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Taldycupes pingiPonomarenko et al. 2014 – Yinping Formation, Anhui, China, Middle Permian (Capitanian)
  • Taldycupes reticulatusPonomarenko, 1969 – Akkolka Formation, Kazakhstan, Late Permian (Changhsingian)
  • Taldycupes rosanoviPonomarenko, 2006 – Limptekon Formation, Krasnoyarsk, Russia, Late Permian (Changhsingian)
  • Taldycupes vasjuchitshevi(Rohdendorf, 1961) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
• Tecticupes Rohdendorf, 1961 (syn StegocupesRohdendorf, 1961)
  • Tecticupes heckeriRohdendorf, 1961 (Type species) – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian) Mitina Formation, Kemerovo, Russia, Middle Permian (Roadian)
  • Tecticupes indistinctusRohdendorf, 1961 – Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian)
  • Tecticupes martynoviPonomarenko, 2013 – Amanak Formation. Orenburg, Middle Permian (Capitanian)
• Tychticupes Rohdendorf, 1961 – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Tychticupes beljanini(Rohdendorf, 1961)
  • Tychticupes radtshenkoiRohdendorf, 1961 (Type species)
  • Tychticupes ragoziniRohdendorf 1961
  • "Tychticupes" ussoviRohdendorf 1961 (nomen dubium)
• Tychticupoides Rohdendorf, 1961 – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Tychticupoides grjazeviRohdendorf, 1961 (Type species)
• Uskaticupes Rohdendorf, 1961 (nomen dubium) – Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Uskaticupes javorskyiRohdendorf, 1961 (nomen dubium)

The genera Alveolacupes Lin, 1983, Clathropenna Fujiyama, 1973 and Penecupes Ren, 1995 were previously placed in the family by some authors, but were all excluded from it by Kirejtshuk (2020). The genera Wuchangia Hong, 1985 and Yuxianocoleus Hong, 1985, both from the Lower Jurassic of Hubei, China, were also excluded from the family by Zhao et al. (2021) (as the subfamily Taldycupedinae). ^[2]

References

1. Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi: 10.3390/geosciences10020073 . ISSN   2076-3263.
2. Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10 e72692. doi: 10.7554/eLife.72692 . ISSN   2050-084X. PMC   8585485 . PMID   34747694.