Thaumastosaurus Temporal range: Late Eocene, | |
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"Mummified" specimens of T. gezei | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Amphibia |
Order: | Anura |
Family: | Pyxicephalidae |
Subfamily: | Pyxicephalinae |
Genus: | † Thaumastosaurus de Stefano, 1903 |
Species | |
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Synonyms | |
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Thaumastosaurus ("marvelous lizard") is an extinct genus of frogs in the family Pyxicephalidae. Five species are known, all from the Eocene of western Europe, including France, England, and Switzerland. Specimens of the species T. gezei from the Quercy Phosphorites of France are known for their exceptional preservation, providing three-dimensional images of the animal's life appearance. [1]
"Mummified" Thaumastosaurus specimens were first described in the 19th century, named " Rana " plicata (a name later found to be preoccupied, and later reclassified as T. gezei) and "Bufo " serratus (later amended to servatus). Fragmentary remains of a skull described in 1903 were confused for the anterior vertebra of a highly unusual lizard, hence the name Thaumastosaurus. Later analyses found "Thaumastosaurus" to be a frog, and assigned the mummified specimens to it. Other skeletal remains assigned to the genus were later found elsewhere in Europe. [1] [2] [3]
Initially, based on comparative anatomy, Thaumastosaurus was classified within the superfamily Hyloidea, although this would be evolutionary improbable as the Hyloidea were largely restricted to the Americas at the time. More recent cranial analyses have found it to belong to the Pyxicephalidae, a family that contains several large frogs that are now found only in Africa, and specifically a stem-member of the subfamily Pyxicephalinae. This suggests that the ancestors of Thaumastosaurus evolved in Africa and dispersed to western Europe, then a tropical island group, during the early Paleogene. It may be related to Rocekophryne , a fossil frog known from the Eocene of Algeria. Thaumastosaurus appears to have gone extinct at the end of the Eocene, during the Eocene–Oligocene extinction event as the climate rapidly cooled. Afterwards, it was replaced by the modern ranid frog genus Pelophylax , which colonized Europe from Asia. [1] [2] [4] [5]
Five species are known, although some may be conspecific with one another:
The Neobatrachia are a suborder of the Anura, the order of frogs and toads.
Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene, spread to Europe by the late Eocene, and further spread to Asia and Africa by the early Miocene. They had largely disappeared worldwide by the late Miocene, with the latest recorded species at the end of the Miocene in Africa. They were among the first carnivorans to evolve large body size. Amphicyonids are colloquially referred to as "bear-dogs".
Palaeobatrachus is an extinct genus of frogs from Europe that existed from the middle Eocene to the middle Pleistocene, spanning almost 50 million years. They were obligately aquatic, and would have not spent much time on dry land. They are one of two genera and by far the largest genus in the family Palaeobatrachidae, which are considered to be members of Pipimorpha, related to the South American-African family Pipidae, which includes the African clawed frog and Surinam toad.
Strigogyps is an extinct genus of prehistoric bird from the Middle Eocene to Early Oligocene of France and Germany. It was probably around the size of a large chicken or a guan, weighing not quite 1 kilogram (2.2 lb). Apparently, as indicated by the ratio of lengths of wing to leg bones, S. sapea was flightless. Its legs were not adapted to running, so it seems to have had a walking lifestyle similar to trumpeters. Unlike other Cariamiformes, which appear to have been mostly carnivorous, Strigogyps specimens suggest a herbivorous diet.
Necromanis is an extinct genus of pangolin from superfamily Manoidea. It lived from the middle Oligocene to middle Miocene in Europe. It was originally placed within family Manidae, but was eventually removed from it as more fossil pholidotids from outside that family were found and studied more extensively. Currently, Necromanis is placed as incertae sedis within the pholidotid superfamily Manoidea, together with the families Manidae and Patriomanidae.
Palaeotheriidae is an extinct family of herbivorous perissodactyl mammals that inhabited Europe, with less abundant remains also known from Asia, from the mid-Eocene to the early Oligocene. They are classified in Equoidea, along with the living family Equidae.
Ursoidea is a superfamily of arctoid carnivoran mammals that includes the families Subparictidae, Amphicynodontidae, and Ursidae which the last family includes the extant lineages of bears as well the extinct Hemicyoninae and Ursavinae. The interrelationships of ursoids has had slight arrangements. In the past it was thought the extinct Amphicyonidae were stem-bears based on morphological analysis of the ear region, though the most recent publications on early amphicyonids suggests they were basal caniforms. The amphicynodontids are sometimes classified as either a subfamily of bears, a paraphyletic assemblage of early bears, or even stem-pinnipeds. The subparictids were previously classified as amphicynodontine/ids. The hemicyonines have been occasionally reclassified as a separate family.
Hyloidea is a superfamily of frogs. Hyloidea accounts for 54% of all living anuran species. The superfamily Hyloidea branched off from its closest relative, the Australobatrachia, during the mid-Cretaceous. The fossil evidence found during the Cretaceous-Paleogene extinction event could not determine the effects upon the frogs, due to the lack of fossils. Increased forestation erupted after this extinction, possibly leading to more arboreal adaptations of these anurans to be best suited for this habitat.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2002.
Lazarussuchus is an extinct genus of amphibious reptile, known from the Cenozoic of Europe. It is the youngest known member of Choristodera, an extinct order of aquatic reptiles that first appeared in the Middle Jurassic. Fossils have been found in Late Paleocene, Late Oligocene, Early Miocene and Late Miocene deposits in France, Germany, and the Czech Republic. Two species have been named: the type species L. inexpectatus ("unexpected") from the late Oligocene of France. and L. dvoraki from the early Miocene of the Czech Republic. It was not a large animal; with the total preserved body and tail length of L. inexpectatus being just over 30 centimetres. A complete specimen of Lazarussuchus with preserved soft tissue was found from the Late Paleocene of France, but has not been assigned to a species.
Mesophryne beipiaoensis is an extinct species of frog, from the Cretaceous Yixian Formation of Liaoning (China), and the only species in the genus Mesophryne. It is known from a single specimen collected near Heitizigou, 25 kilometres (16 mi) south of Beipiao, from which the specific epithet derives. The specimen has a snout–vent length of 69 millimetres (2.7 in). While some authors have suggested Mesophryne is a synonym of Liaobatrachus, this has been rejected by other authors. In a phylogenetic analysis it was found to be a crown group frog, which was more derived than Ascaphus and Leiopelma, but less so than alytids and other more advanced frogs.
Eopelobates is an extinct genus of frogs in the family Pelobatidae. Closely related to the living European spadefoot toad, it is known from the Eocene of western North America, and the Eocene–Pliocene of Europe. It is suggested that the distribution over both Europe and North America is due to dispersal during the Paleocene-Eocene Thermal Maximum.
Albionbatrachus is an extinct genus of prehistoric frogs from England and Romania. Two species are recognized: While previously synonymised with Palaeobatrachus, it is now considered a distinct palaeobatrachid genus based on characters of the frontoparietals.
Liaobatrachus is a genus of prehistoric frog, the first fossil specimen of which was recovered from the Yixian Formation of Liaoning Province, China. It was the first Mesozoic era frog ever found in China. The species Callobatrachus sanyanensis,Mesophryne beipiaoensis and Yizhoubatrachusmacilentus were classified as species of Liaobatrachus in one study, but this has been rejected by other authors. The genus has been considered a nomen dubium by some authors due to the poor preservation of the holotype specimen. Fossils were found in the Sihetun locality of the western part of Liaoning province, in the lower part of the Yixian Formation, and date to approximately 124.6 Ma. Another specimen was collected near Heitizigou, 25 kilometres (16 mi) south of Beipiao. The specimen has a snout–vent length of 69 millimetres (2.7 in). Liaobatrachus is considered to be the most basal member of Discoglossidae based on phylogenetic analysis.
Vulcanobatrachus is an extinct genus of fossil frog. The genus contains the single species Vulcanobatrachus mandelai found at Marydale, South Africa, described in 2005 and named after Nelson Mandela. The genus owes its name to the fact that fossils were recovered from an extinct volcanic crater lake of Late Cretaceous age. The fossil frogs are assumed to have died following a limnic eruption (a degassing event possibly of CO2) by the volcano.
The Pyxicephalidae are a family of frogs currently found in sub-Saharan Africa. However, in the Eocene, the taxon Thaumastosaurus lived in Europe.
Asiavorator is an extinct genus of civet-like carnivoran belonging in the family Stenoplesictidae. It was endemic to Asia and lived during the Eocene and Oligocene epochs.
Pipoidea are a clade of frogs, that contains the most recent common ancestor of living Pipidae and Rhinophrynidae as well as all its descendants. It is broadly equivalent to Xenoanura.
The Quercy Phosphorites Formation is a geologic formation and lagerstätte in Occitanie, southern France. It preserves fossils dated to the Paleogene period, or MP16 to MP28 zones of the European land mammal age classification, ranging from approximately 38 to 25 Ma.
Palaeobatrachidae is an extinct family of frogs known from the Late Cretaceous to the Pleistocene primarily of Europe. They were highly adapted to aquatic life, like other members of the Pipimorpha. The oldest undoubted records of the family are from the lower Campanian of France. By far the most abundant genus is Palaeobatrachus, known from the Eocene to Pleistocene of Europe, with most other named genera in the family synonymised with it, the only exception being Albionbatrachus, which is distinguished from Palaeobatrachus by characters of its frontoparietals. The youngest fossils of Palaeobatrachus date to around 500,000 years ago, during the Middle Pleistocene after which they likely became extinct due to increasing aridity and freezing temperatures during the ice ages. Fossils of indeterminate palaeobatrachids are also known from the Pliocene and Early Pleistocene of Western Siberia.