Trinodus

Last updated

Trinodus
Temporal range: Ordovician
Arthrorhachis tarda draw entite.png
A drawing of Trinodus tardus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Trilobita (?)
Order: Agnostida
Family: Metagnostidae
Genus: Trinodus
M'Coy, 1846
Type species
Trinodus agnostiformes
M'Coy, 1846
Species
  • Trinodus agnostiformesM'Coy, 1846 synonyms A. agnostiformis, A. trinodus
  • Trinodus tardus(Barrande,1846) synonyms Battus tardus, Agnostus glabratus, A. tardus [1] Arthrorhachis tarda, alternative spelling A. tardus
  • Trinodus elspethi(Raymond, 1925) synonym Arthrorhachis elspethi
  • Trinodus danicusPoulsen, 1965 synonym Arthrorhachis danica
  • Trinodus girvanensis(Reed, 1903) synonyms Agnostus girvanensis, Arthrorhachis girvanensis, Girvanagnostus girvanensis
  • Trinodus hupehensisLu, 1975 synonym Arthrorhachis hupehensis
  • Trinodus knockerkensisRomano & Owen, 1993 synonym Arthrorhachis knockerkensis
  • Trinodus latilimbata(Ju in Qiu et al., 1983)synonym Arthrorhachis latilimbata
  • Trinodus pragensis(Přibyl & Vaněk, 1968) synonym Arthrorhachis pragensis
Synonyms
  • Arthrorhachis
  • Girvanagnostus
  • Metagnostus

Trinodus is a very small to small (about 1 centimetre or 0.39 inches) blind trilobite, a well known group of extinct marine arthropods, which lived during the Ordovician (Tremadocian to early Hirnantian), [1] in what are now the Yukon Territories, Virginia, Italy, Czech Republic, Poland, Denmark, Sweden, Svalbard, Ireland, Scotland, Wales, Iran, Kazakhstan and China. It is one of the last of the Agnostida order to survive.

Contents

Etymology

Trinodus is derived from the Latin tri (three) and nodus (node).
Arthrorhachis is derived from the Greek ἄρθρον (árthron, "joint") and ῥάχις (rháchis, meaning axis, spine, ridge or backbone). [1]

Taxonomy

Trinodus, Arthrorhachis and Geragnostus are closely related and it may be appropriate to assign their species to just one genus. All species in these three genera have virtually identical cephalons, but of T. agnostiformes, the type species of Trinodus only one poorly preserved cephalon was known. However, relatively recent, pygidia assignable to T. agnostiformes were found. Although this material is distorted or incompletely preserved, it is very similar to the pygidium of Arthrorhachis tarda. Species with a rear rhachis lobe longer than the postaxial region are henceforth combined in Geragnostus, all others are assigned to Trinodus. [2]

Species previously assigned to Trinodus

Distribution

Development

Trinodus elspethi, which - as an agnostoid - only has two thorax segments, has at least nine larval stages (or instars), three meraspid and six holaspid, in its life. So it molted at least eight times. [18]

Related Research Articles

<span class="mw-page-title-main">Ordovician</span> Second period of the Paleozoic Era 485–444 million years ago

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Ma) to the start of the Silurian Period 443.8 Mya.

<span class="mw-page-title-main">John William Salter</span> English naturalist, geologist, and palaeontologist

John William Salter was an English naturalist, geologist, and palaeontologist.

The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 1.4 million years, from 445.2 to 443.8 Ma. The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation.

<i>Han <span style="font-style:normal;">(trilobite)</span></i> Extinct genus of trilobites

Han is a monotypic genus of agnostid trilobite, whose sole member is Han solo. The type specimen of H. solo was found in marine strata of the Arenig to Llanvirn-aged Zitai Formation of Middle Ordovician southern China, and is named after the character in Star Wars.

<i>Agnostus</i> Extinct genus of trilobites

Agnostus is a genus of agnostid trilobites, belonging to the family Agnostidae, that lived during the late Middle Cambrian – early Upper Cambrian. It is the type genus of the family Agnostidae and is subdivided into two subgenera, Agnostus and Homagnostus.

<i>Flexicalymene</i> Genus of trilobites (fossil)

Flexicalymene Shirley, 1936. is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and Family Calymenidae. Flexicalymene specimens can be mistaken for Calymene, Gravicalymene, Diacalymene and a few other Calymenina genera. They are used as an index fossil in the Ordovician. Ohio and North America are particularly known for being rich with Flexicalymene fossils.

<i>Gravicalymene</i>

Gravicalymene Shirley, 1936, is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and family Calymenidae. Species included in this genus have previously been allocated to Calymene Brongniart 1822,Flexicalymene Shirley, 1936. and Sthenarocalymene Siveter 1977.

<span class="mw-page-title-main">Arenig</span> Time interval during the Ordovician period

In geology, the Arenig is a time interval during the Ordovician period and also the suite of rocks which were deposited during this interval.

<span class="mw-page-title-main">Katian</span>

The Katian is the second stage of the Upper Ordovician. It is preceded by the Sandbian and succeeded by the Hirnantian Stage. The Katian began 453 million years ago and lasted for about 7.8 million years until the beginning of the Hirnantian 445.2 million years ago. During the Katian the climate cooled which started the Late Ordovician glaciation.

Plumulites is an extinct genus of machaeridians, extinct annelid group.

<span class="mw-page-title-main">Soom Shale</span> Late Ordovician lagerstatte formation in South Africa

The Soom Shale is a member of the Late Ordovician (Hirnantian) Cederberg Formation in South Africa, renowned for its remarkable preservation of soft-tissue in fossil material. Deposited in still waters, the unit lacks bioturbation, perhaps indicating anoxic conditions.

<i>Sphaeragnostus</i>

Sphaeragnostus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It can be recognized by having two thorax segments, a totally effaced headshield, while the tailshield although effaced, has a clear furrow parallel to its border, and a short, convex, subcircular axis. It lived during the Ordovician.

<i>Pleuroctenium</i>

Pleuroctenium Hawle & Corda (1847) is an agnostid trilobite belonging to the family Condylopygidae Raymond (1913). The genus occurs in Middle Cambrian (Drumian) strata of Canada, the Czech Republic, England and Wales, France, and Sweden.

<i>Geragnostus</i> Extinct genus of trilobites

Geragnostus is a genus of very small agnostid trilobites whose fossils are found Ordovician-aged marine strata from Eurasia, North America and Argentina.

Sagavia is a genus of trilobites that lived during the Middle and Upper Ordovician in what are now Northwest and Southeast China, North Kazakhstan and Wales. It is a typical cyclopygid that can be distinguished by its large but separate eyes, elongated glabella, five thorax segments and a pygidium with clearly defined axis and border.

<span class="mw-page-title-main">Trimerellida</span> Extinct order of brachiopods

Trimerellida is an extinct order of craniate brachiopods, containing the sole superfamily Trimerelloidea and the families Adensuidae, Trimerellidae, and Ussuniidae. Trimerellidae was a widespread family of warm-water brachiopods ranging from the Middle Ordovician to the late Silurian (Ludlow). Adensuidae and Ussuniidae are monogeneric families restricted to the Ordovician of Kazakhstan. Most individuals were free-living, though some clustered into large congregations similar to modern oyster reefs.

<i>Gog</i> (trilobite) Extinct genus of trilobites

Gog is a genus of large, flattened asaphid trilobite from the Middle Arenig-aged Svalbard, Valhallfonna Formation, Olenidsletta, Member, of Spitzbergen, Norway, and the Upper Arenig-aged Dawan Formation in Hubei, China.

<span class="mw-page-title-main">Siphonotretida</span> Extinct order of marine lamp shells

Siphonotretida is an extinct order of linguliform brachiopods in the class Lingulata. The order is equivalent to the sole superfamily Siphonotretoidea, itself containing the sole family Siphonotretidae. Siphonotretoids were originally named as a superfamily of Acrotretida, before being raised to their own order.

<span class="mw-page-title-main">Nod Glas Formation</span>

The Nod Glas Formation is an Ordovician lithostratigraphic group in Mid Wales. The rock of the formation is made up of pyritous, graptolitic mudstone that is generally black in colour. It weathers to a soft, very well cleaved and coal-like material. The formation runs from Conwy in the north, down to Cardigan Bay in the area around Aberdyfi and Tywyn, though it is not a continuous over this area.

<span class="mw-page-title-main">Saint-Chinian Formation</span>

The Saint-Chinian Formation is a geological formation composed of shales with limestone inclusions, dating from the Lower Ordovician (Tremadocian).

References

  1. 1 2 3 4 5 Budil, P.; Fatka, O.; Kolář, P.; David, M. (2011). "Arthrorhachis Hawle & Corda, 1847 (Agnostida) in the Prague Basin revisited" (PDF). Bulletin of Geosciences. 86 (4): 707–724. doi: 10.3140/bull.geosci.1262 .
  2. 1 2 Turvey, S.T. (2005). "Agnostid trilobites from the Arenig-Llanvirn of South China". Transactions of the Royal Society of Edinburgh: Earth Sciences. 95 (3–4): 527–542. doi:10.1017/s026359330000119x. S2CID   130775617.
  3. 1 2 Karim, T.S. (2009). "Ordovician trilobites from Iran". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 99 (2): 101–124. doi:10.1017/s1755691009007063. S2CID   129780250.
  4. Fortey, R. A. (1980). "The Ordovician trilobites of Spitsbergen. III. Remaining trilobites of the Valhallfonna Formation". Norsk Polarinstitutt Skrifter. 171: 1–163.
  5. Romano, M.; Owen, A. W. (1993). "Early Caradoc Trilobites of Eastern Ireland and their paleogeographical significance". Palaeontology. 36 (3): 681–720.
  6. 1 2 3 4 Ahlberg, P. (1989). "Agnostid trilobites from the Upper Ordovician of Sweden and Bornholm, Denmark" (PDF). Bulletin of the Geological Society of Denmark. 37: 213–226. doi:10.37570/bgsd-1988-37-17.
  7. Chlupac, I.; Havlicek, V.; Kukal, Z.; Storch, P. (1998). Palaeozoic of the Barrandian (Cambrian to Devonian). pp. 1–183.
  8. Leone, F.; Hammann, W.; Laske, R.; Serpagli, E.; Villas, E. (1991). "Lithostratigraphic units and biostratigraphy of the post-sardic Ordovician sequence in south-west Sardinia". Bollettino della Societá Paleontologica Italiana. 30: 201–235.
  9. Havlicek, V.; Vanek, J. (1990). "Ordovician Invertebrate communities in black-shale lithofacies (Prague Basin, Czechoslovakia)". Vestnik Ustredniho Ustavu Geologickeho. 65: 223–236.
  10. Ludvigsen, R. (1981). "Biostratigraphical significance of Middle Ordovician trilobites from the Road River Formation, northern Cordillera". Geological Association of Canada, Abstracts. 6 (A-36).
  11. Stratigraphic Group of Yunnan, China (1978). Regional stratigraphic data of Southwest China, Yunnan Province. Beijing: Geological Publishing House.
  12. Temple, J.T. (1965). "Upper Ordovician Brachiopods from Poland and Britain". Acta Palaeontologica Polonica . 10: 379–450.
  13. Price, D. (1981). "Ashgill trilobite faunas from the Llyn Peninsula, North Wales, U.K". Geological Magazine . 16 (3): 201–216. Bibcode:1981GeolJ..16..201P. doi:10.1002/gj.3350160305.
  14. Bassett, D.A.; Whittington, H.B.; Williams, A. (1966). "The Stratigraphy of the Bala District, Merionethshire". Quarterly Journal of the Geological Society of London. 122 (3): 219–269. doi:10.1144/gsjgs.122.1.0219. S2CID   128598649.
  15. Price, D.; Magor, P.M. (1984). "The ecological significance of variation in the genesis composition of Rawtheyan (Late Ordovician) trilobite faunas from North Wales, U.K". Geological Journal. 19 (2): 187–200. Bibcode:1984GeolJ..19..187P. doi:10.1002/gj.3350190207.
  16. Fortey, R. A.; Owens, R.M. (1978). "Early Ordovician (Arenig) stratigraphy and faunas of the Carmarthen District, South-West Wales". Bulletin of the British Museum (Natural History), Geology Series. 30 (3): 225–294.
  17. Tripp, R.P.; Williams, A.; Paul, C.R.C. (1981). "On an exposure of the Ordovician superstes Mudstones at Colmonell, Gircan District, Strathclyde". Scottish Journal of Geology. 17 (1): 21–25. Bibcode:1981ScJG...17...21T. doi:10.1144/sjg17010021. S2CID   128426864.
  18. Whittington, H. B. et al. Part O, Treatise on Invertebrate Paleontology. Revised, Volume 1 – Trilobita – Introduction, Order Agnostida, Order Redlichiida. 1997