Ukureyskaya Formation | |
---|---|
Stratigraphic range: | |
Type | Geological formation |
Sub-units | Lower, Middle and Upper subformations |
Underlies | Utanskaya Formation |
Overlies | Kulindinskaya Formation |
Thickness | Several hundred metres |
Lithology | |
Primary | Sandstone, siltstone, tuffite |
Other | Mudstone |
Location | |
Coordinates | 52°30′N116°42′E / 52.5°N 116.7°E |
Approximate paleocoordinates | 58°00′N117°12′E / 58.0°N 117.2°E |
Region | Zabaykalsky Krai |
Country | Russia |
Extent | Zabaykalsky Krai |
The Ukureyskaya Formation, also referred to as the Ukurey Formation, is a geological formation in Zabaykalsky Krai, part of the Russian Far East. It is made up of Middle Jurassic and Late Jurassic layers. [1] It covers large areas around Kulinda. [2]
The Ukureyskaya Formation was mentioned by Sinitsa & Starukhina (1986), [3] and was first discovered by Sofia M. Sinitsa and her team from the Russian Academy of Sciences before being excavated in 2010 by a group of Russian and Belgian palaeontologists; [4] during this time, the holotype of Kulindadromeus zabaikalicus was discovered. [5] Excavations wrapped up in 2013 or 2014. [6]
The volcanic ash layers of the Ukureyskaya Formation form a Konservat-Lagerstätte with an exceptional preservation, [1] and it likely represents a nearshore lacustrine or estuarine-deltaic environment similar to the Yixian Formation. [2]
In total, two bone beds were identified during the excavations that took place between 2010 and 2013/14; [6] Bonebed 4 is older than Bonebed 3.
Sinitsa & Starukhina (1986) and Sinitsa (2011) suggested that the Ukureyskaya Formation dated to the Late Jurassic-Early Cretaceous. [3] [4]
Godefroit et al. (2014) and Alivanov & Saveliev (2014) have suggested that as a whole, the Ukureyskaya Formation dates to the Bajocian-Tithonian, [5] [7] while more recent dating work by Cincotta et al. (2019) suggests that the layers containing the remains of Kulindadromeus are Bathonian in age. [8]
Genus | Species | Location | Stratigraphic position | Material | Notes | Image |
---|---|---|---|---|---|---|
Crustacea [2] | Indeterminate [2] | Kulinda [2] | "Indeterminate remains" [2] | |||
Insecta [5] | Indeterminate [5] | Kulinda [5] | Represented by multiple species. [5] | |||
Kulindadromeus' [5] [7] | K. zabaikalicus [5] [7] | Kulinda [5] | "Base of the Ukureyska Formation" [5] | "Hundreds of disarticulated skeletons including six skulls" [5] | Synonyms include Daurosaurus olovus and Lepidocheirosaurus natalis. [9] | |
Ornithopoda [2] | Indeterminate [2] | Kulinda [2] | "Indeterminate remains" [2] | Represents an unnamed genus that is distinct from Kulindadromeus . [2] | ||
Theropoda [2] | Indeterminate [2] | Kulinda [2] | "Single tooth" [2] | Based on undiagnostic remains. [2] |
Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group, as birds are theropod dinosaurs.
Ornithomimosauria are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia, as well as Africa and possibly Australia. The group first appeared in the Early Cretaceous and persisted until the Late Cretaceous. Primitive members of the group include Nqwebasaurus, Pelecanimimus, Shenzhousaurus, Hexing and Deinocheirus, the arms of which reached 2.4 m (8 feet) in length. More advanced species, members of the family Ornithomimidae, include Gallimimus, Struthiomimus, and Ornithomimus. Some paleontologists, like Paul Sereno, consider the enigmatic alvarezsaurids to be close relatives of the ornithomimosaurs and place them together in the superfamily Ornithomimoidea.
Dryosaurus is a genus of an ornithopod dinosaur that lived in the Late Jurassic period. It was an iguanodont. Fossils have been found in the western United States and were first discovered in the late 19th century. Valdosaurus canaliculatus and Dysalotosaurus lettowvorbecki were both formerly considered to represent species of Dryosaurus.
Camptosaurus is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard'.
Yixianosaurus is a maniraptoran theropod dinosaur genus from the Early Cretaceous of China.
Neornithischia is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs.
Laosaurus is a genus of neornithischian dinosaur. The type species, Laosaurus celer, was first described by O.C. Marsh in 1878 from remains from the Oxfordian-Tithonian-age Upper Jurassic Morrison Formation of Wyoming. The validity of this genus is doubtful because it is based on fragmentary fossils. A second species from the Morrison Formation, L. gracilis, and a species from the late Cretaceous Allison Formation of Alberta, Canada, Laosaurus minimus, are also considered dubious.
Lycorhinus is a genus of heterodontosaurid ornithischian dinosaur from the Early Jurassic strata of the Elliot Formation located in the Cape Province, South Africa.
Wulagasaurus is a genus of saurolophine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China.
Avifilopluma is a clade containing all animals with feathers. Unlike most clades, which are defined based on relative relationships, Avifilopluma is defined based on an apomorphy, that is, a unique physical characteristic shared by one group and not found outside that group. Its content is unclear, and has been speculated to range from Coelurosauria to all of Ornithodira.
Fruitadens is a genus of heterodontosaurid dinosaur. The name means "Fruita teeth", in reference to Fruita, Colorado (USA), where its fossils were first found. It is known from partial skulls and skeletons from at least four individuals of differing biological ages, found in Tithonian rocks of the Morrison Formation in Colorado. Fruitadens is one of the smallest known ornithischian dinosaur, with young adults estimated at 65 to 75 cm in length and 0.5 to 0.75 kg in weight. It is interpreted as an omnivore and represents one of the latest-surviving heterodontosaurids.
Arkharavia is a dubious genus of somphospondylan sauropod, but at least some of the remains probably belong to a hadrosaurid. It was discovered in the Udurchukan Formation in Russia and lived during the Late Cretaceous. It was described in 2010 by Alifanov and Bolotsky and the type species is A. heterocoelica.
Koreanosaurus is a genus of orodromine neornithischian dinosaur. One species has been described, Koreanosaurus boseongensis.
Chernyshevsky District is an administrative and municipal district (raion), one of the thirty-one in Zabaykalsky Krai, Russia. It is located in the center of the krai and borders Tungokochensky District in the north, Sretensky District in the east, District in the south, and Nerchinsky District in the west. The area of the district is 13,200 square kilometers (5,100 sq mi). Its administrative center is the urban locality of Chernyshevsk. As of the 2010 Census, the total population of the district was 35,019, with the population of Chernyshevsk accounting for 38.2% of that number.
Kulindadromeus was a herbivorous dinosaur, a basal neornithischian from the Middle Jurassic. The first Kulindadromeus fossil was found in Russia. Its feather-like integument is evidence for protofeathers being basal to Ornithischia and possibly Dinosauria as a whole, rather than just to Coelurosauria, as previously suspected.
Laquintasaura is a genus of Venezuelan ornithischian dinosaur containing only the type species Laquintasaura venezuelae. It is known for being one of the and most primitive ornithischians in the fossil record, as well as the first dinosaur to have been identified from Venezuela. The name is derived from the La Quinta Formation, where it was discovered and the feminine Greek suffix for lizard, with the specific name referring to the country of Venezuela. It is known from hundreds of fossil elements, all derived from a single extensive bonebed locality. Initially discovered by French palaeontologists, numerous expeditions have been conducted to excavate from the bonebed, largely led by Marcelo R Sánchez-Villagra. Once thought to represent remains of Lesothosaurus, it was formally named in a 2014 study; much of the abundant material was not yet prepared at the time and research remains ongoing.
This timeline of ornithomimosaur research is a chronological listing of events in the history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Although fragmentary, probable, ornithomimosaur fossils had been described as far back as the 1860s, the first ornithomimosaur to be recognized as belonging to a new family distinct from other theropods was Ornithomimus velox, described by Othniel Charles Marsh in 1890. Thus the ornithomimid ornithomimosaurs were one of the first major Mesozoic theropod groups to be recognized in the fossil record. The description of a second ornithomimosaur genus did not happen until nearly 30 years later, when Henry Fairfield Osborn described Struthiomimus in 1917. Later in the 20th century, significant ornithomimosaur discoveries began occurring in Asia. The first was a bonebed of "Ornithomimus" asiaticus found at Iren Debasu. More Asian discoveries took place even later in the 20th century, including the disembodied arms of Deinocheirus mirificus and the new genus Gallimimus bullatus. The formal naming of the Ornithomimosauria itself was performed by Rinchen Barsbold in 1976.
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