Ukureyskaya Formation | |
---|---|
Stratigraphic range: | |
Type | Geological formation |
Sub-units | Lower, Middle and Upper subformations |
Underlies | Utanskaya Formation |
Overlies | Kulindinskaya Formation |
Thickness | Several hundred metres |
Lithology | |
Primary | Sandstone, siltstone, tuffite |
Other | Mudstone |
Location | |
Coordinates | 52°30′N116°42′E / 52.5°N 116.7°E |
Approximate paleocoordinates | 58°00′N117°12′E / 58.0°N 117.2°E |
Region | Zabaykalsky Krai |
Country | Russia |
Extent | Zabaykalsky Krai |
The Ukureyskaya Formation, also referred to as the Ukurey Formation, is a geological formation in Zabaykalsky Krai, part of the Russian Far East. It is made up of Middle Jurassic and Late Jurassic layers. [1] It covers large areas around Kulinda. [2]
The Ukureyskaya Formation was mentioned by Sinitsa & Starukhina (1986), [3] and was first discovered by Sofia M. Sinitsa and her team from the Russian Academy of Sciences before being excavated in 2010 by a group of Russian and Belgian palaeontologists; [4] during this time, the holotype of Kulindadromeus zabaikalicus was discovered. [5] Excavations wrapped up in 2013 or 2014. [6]
The volcanic ash layers of the Ukureyskaya Formation form a Konservat-Lagerstätte with an exceptional preservation, [1] and it likely represents a nearshore lacustrine or estuarine-deltaic environment similar to the Yixian Formation. [2]
In total, two bone beds were identified during the excavations that took place between 2010 and 2013/14; [6] Bonebed 4 is older than Bonebed 3.
Sinitsa & Starukhina (1986) and Sinitsa (2011) suggested that the Ukureyskaya Formation dated to the Late Jurassic-Early Cretaceous. [3] [4]
Godefroit et al. (2014) and Alivanov & Saveliev (2014) have suggested that as a whole, the Ukureyskaya Formation dates to the Bajocian-Tithonian, [5] [7] while more recent dating work by Cincotta et al. (2019) suggests that the layers containing the remains of Kulindadromeus are Bathonian in age. [8]
Genus | Species | Location | Stratigraphic position | Material | Notes | Image |
---|---|---|---|---|---|---|
Crustacea [2] | Indeterminate [2] | Kulinda [2] | "Indeterminate remains" [2] | |||
Insecta [5] | Indeterminate [5] | Kulinda [5] | Represented by multiple species. [5] | ![]() | ||
Kulindadromeus [5] [7] | K. zabaikalicus [5] [7] | Kulinda [5] | "Base of the Ukureyska Formation" [5] | "Hundreds of disarticulated skeletons including six skulls" [5] | Synonyms include Daurosaurus olovus and Lepidocheirosaurus natalis. [9] | ![]() |
Ornithopoda [2] | Indeterminate [2] | Kulinda [2] | "Indeterminate remains" [2] | Represents an unnamed genus that is distinct from Kulindadromeus . [2] | ||
Theropoda [2] | Indeterminate [2] | Kulinda [2] | "Single tooth" [2] | Based on undiagnostic remains. [2] |
Ornithischia is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name Ornithischia, or "bird-hipped", reflects this similarity and is derived from the Greek stem ornith- (ὀρνιθ-), meaning "bird", and ischion (ἴσχιον), meaning "hip". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs, the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs (Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous pelts, and there is much debate over whether these filaments found in specimens of Tianyulong, Psittacosaurus, and Kulindadromeus may have been primitive feathers.
A feathered dinosaur is any species of dinosaur possessing feathers. That includes all species of birds, and in recent decades evidence has accumulated that many non-avian dinosaur species also possessed feathers in some shape or form. The extent to which feathers or feather-like structures were present in dinosaurs as a whole is a subject of ongoing debate and research.
Ornithomimosauria are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia, as well as Africa and possibly Australia. The group first appeared in the Early Cretaceous and persisted until the Late Cretaceous. Primitive members of the group include Nqwebasaurus, Pelecanimimus, Shenzhousaurus, Hexing and Deinocheirus, the arms of which reached 2.4 m (8 feet) in length. More advanced species, members of the family Ornithomimidae, include Gallimimus, Struthiomimus, and Ornithomimus. Some paleontologists, like Paul Sereno, consider the enigmatic alvarezsaurids to be close relatives of the ornithomimosaurs and place them together in the superfamily Ornithomimoidea.
Camptosaurus is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard'.
Yixianosaurus is a maniraptoran theropod dinosaur genus from the Early Cretaceous of China.
Bissektipelta is a genus of ankylosaurine thyreophoran dinosaurs that lived in Asia during the Late Cretaceous in what is now the Bissekty Formation of Uzbekistan. Bissektipelta is a monospecific genus, containing only the type species B. archibaldi.
Neornithischia is a clade of the dinosaur order Ornithischia. It is the sister group of the Thyreophora within the clade Genasauria. Neornithischians are united by having a thicker layer of asymmetrical enamel on the inside of their lower teeth. The teeth wore unevenly with chewing and developed sharp ridges that allowed neornithischians to break down tougher plant food than other dinosaurs. Neornithischians include a variety of basal forms historically known as "hypsilophodonts", including the Parksosauridae; in addition, there are derived forms classified in the groups Marginocephalia and Ornithopoda. The former includes clades Pachycephalosauria and Ceratopsia, while the latter typically includes Hypsilophodon and the more derived Iguanodontia.
Lycorhinus is a genus of heterodontosaurid ornithischian dinosaur from the Early Jurassic strata of the Elliot Formation located in the Cape Province, South Africa.
Wulagasaurus is a genus of saurolophine hadrosaurid dinosaur from the Late Cretaceous of Heilongjiang, China.
Avifilopluma is a clade containing all animals with feathers. Unlike most clades, which are defined based on relative relationships, Avifilopluma is defined based on an apomorphy, that is, a unique physical characteristic shared by one group and not found outside that group. Its content is unclear, and has been speculated to range from Coelurosauria to all of Ornithodira.
Fruitadens is a genus of heterodontosaurid dinosaur. The name means "Fruita teeth", in reference to Fruita, Colorado (USA), where its fossils were first found. It is known from partial skulls and skeletons from at least four individuals of differing biological ages, found in Tithonian rocks of the Morrison Formation in Colorado. Fruitadens is one of the smallest known ornithischian dinosaur, with young adults estimated at 65 to 75 cm in length and 0.5 to 0.75 kg in weight. It is interpreted as an omnivore and represents one of the latest-surviving heterodontosaurids.
Arkharavia is a dubious genus of somphospondylan sauropod, but at least some of the remains probably belong to a hadrosaurid. It was discovered in the Udurchukan Formation in Russia and lived during the Late Cretaceous. It was described in 2010 by Alifanov and Bolotsky and the type species is A. heterocoelica.
Koreanosaurus is a genus of orodromine neornithischian dinosaur. One species has been described, Koreanosaurus boseongensis.
Kulindadromeus was a herbivorous dinosaur, a basal neornithischian from the Middle Jurassic. The first Kulindadromeus fossil was found in Russia. Its feather-like integument is evidence for protofeathers being basal to Ornithischia and possibly Dinosauria as a whole, rather than just to Coelurosauria, as previously suspected.
Laquintasaura is a genus of Venezuelan ornithischian dinosaur containing only the type species Laquintasaura venezuelae. The species was the first dinosaur to have been identified from Venezuela. It is known from extensive remains, all from a single bonebed locality which has been sampled for specimen blocks over the course of several expeditions, largely led by Marcelo R Sánchez-Villagra. A small animal, it is known for its distinct dental anatomy and for being one of the earliest and most primitive ornithischians in the fossil record. Taxonomic uncertainty has led to conflicting hypotheses that it is either as the base of Ornithischia or at the base of the subgroup Thyreophora. In either model, its dating to around 200 million years ago at the start of the Jurassic, existence in equatorial latitudes, and primitive nature make it a key insight into early ornithischian evolution. It is thought that Laquintasaura would have lived in groups and had a possible omnivorous diet, living on a seasonal alluvial plain and being preyed upon by the contemporary Tachiraptor.
This timeline of ornithomimosaur research is a chronological listing of events in the history of paleontology focused on the ornithomimosaurs, a group of bird-like theropods popularly known as the ostrich dinosaurs. Although fragmentary, probable, ornithomimosaur fossils had been described as far back as the 1860s, the first ornithomimosaur to be recognized as belonging to a new family distinct from other theropods was Ornithomimus velox, described by Othniel Charles Marsh in 1890. Thus the ornithomimid ornithomimosaurs were one of the first major Mesozoic theropod groups to be recognized in the fossil record. The description of a second ornithomimosaur genus did not happen until nearly 30 years later, when Henry Fairfield Osborn described Struthiomimus in 1917. Later in the 20th century, significant ornithomimosaur discoveries began occurring in Asia. The first was a bonebed of "Ornithomimus" asiaticus found at Iren Debasu. More Asian discoveries took place even later in the 20th century, including the disembodied arms of Deinocheirus mirificus and the new genus Gallimimus bullatus. The formal naming of the Ornithomimosauria itself was performed by Rinchen Barsbold in 1976.
Anchiornithidae is a family of small paravian dinosaurs. Anchiornithids have been classified at varying positions in the paravian tree, with some scientists classifying them as a distinct family, a basal subfamily of Troodontidae, members of Archaeopterygidae, or an assemblage of dinosaurs that are an evolutionary grade within Avialae or Paraves.
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