| Ursus ingressus | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Carnivora |
| Family: | Ursidae |
| Genus: | Ursus |
| Species: | †U. ingressus |
| Binomial name | |
| †Ursus ingressus Rabeder et al., 2004 [1] | |
Ursus ingressus (the Gamssulzen Cave bear) is an extinct species of the family Ursidae that lived in Central Europe during the Late Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found. [2]
Ursus ingressus was a large cave bear with massive, bulky limbs. It was larger than Ursus spelaeus , which has been estimated to weigh an average of 350 to 600 kg (770 to 1,320 lb) (male specimen). [2] [3]
Some studies have suggested the Gamssulzen Cave bear to have been herbivorous, living off vegetation with little contribution of grass. [4] Other studies proposed Ursus ingressus to have been an omnivore, with participation of terrestrial and more likely aquatic animal protein, that exceeds the participation of animal protein in the diet of the modern brown bear (Ursus arctos). [5] However it has also been suggested, that the feeding habits of cave bears can vary heavily depending on the environment. [6]
Skulls with bite damage from Zoolithen Cave in Germany suggest that Ursus ingressus came into conflicts with other big carnivores of the Late Pleistocene of Europe like the cave lion (Panthera spelaea) or the cave hyena (Crocuta crocuta spelaea). [7]
It has been suggested that the Gamssulzen Cave bear dominated Ursus spelaeus in Central and Eastern Europe, while being out-competed by this species in Western Europe. Around 50,000 years ago, the Gamssulzen Cave bear migrated into the Alps and replaced two former populations of Ursus spelaeus: Ursus spelaeus eremus and Ursus spelaeus ladinicus. Ursus ingressus has been found as far east as the Ural Mountains in Russia and as far west as the Swabian Jura in Germany. [6] [8]
It has mostly been found in medium and high elevated regions and probably was adapted to continental environments with cold and arid climate. [9]
Ursus ingressus and Ursus spelaeus evolved from Ursus deningeri and probably diverged between 173,000 and 414,000, or possibly as much as 600,000 years ago. [6] [2] Some studies still question whether U. ingressus and U. spelaeus are separate species, instead treating them as subspecies of a single species. [9]
The Gamssulzen Cave bear survived U. spelaeus for about 1000 to 2000 years, locally replacing this species, but also became extinct about 30,000 years ago, just prior to the Last Glacial Maximum. The reasons for their extinction are still being discussed, with climate change and human hunting suggested as possible reasons. [8] [10]
Panthera is a genus within the family Felidae, and one of two extant genera in the subfamily Pantherinae. It contains the largest living members of the cat family. There are five living species: the jaguar, leopard, lion, snow leopard and tiger, as well as a number of extinct species, including the cave lion and American lion.
The cave bear is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum.
Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998), and Jin et al. (2007).
Ursus is a genus in the family Ursidae (bears) that includes the widely distributed brown bear, the polar bear, the American black bear, and the Asian black bear. The name is derived from the Latin ursus, meaning bear.
Crocuta is a genus of hyena containing the largest extant member of the family, the spotted hyena (Crocuta crocuta). Several fossil species are also known, with the Pleistocene Eurasian cave hyenas either being regarded as distinct species or subspecies of the spotted hyena.
Panthera spelaea, commonly known as the cave lion, is an extinct Panthera species that was native to Eurasia and northwest North America during the Pleistocene epoch. Genetic analysis of ancient DNA has revealed that while closely related, it was a distinct species genetically isolated from the modern lion, with the genetic divergence between the two species estimated at around 500,000 years ago. The earliest fossils of the P. spelaea lineage in Eurasia date to around 700,000 years ago. It is closely related and probably ancestral to the American lion. The species ranged from Western Europe to eastern Beringia in North America, and was a prominent member of the mammoth steppe fauna, and an important apex predator across its range. It became extinct about 13,000 years ago. It closely resembled living lions with a coat of yellowish-grey fur though unlike extant lions, males appear to have lacked manes.
Pachycrocuta is an extinct genus of prehistoric hyenas. The largest and most well-researched species is Pachycrocuta brevirostris, colloquially known as the giant short-faced hyena as it stood about 90–100 cm (35–39 in) at the shoulder and it is estimated to have averaged 110 kg (240 lb) in weight, approaching the size of a lioness, making it the largest known hyena. Pachycrocuta first appeared during the late Miocene. By 800,000 years ago, it became locally extinct in Europe, with it surviving in East Asia until at least 500,000 years ago, and possibly later elsewhere in Asia.
The history of lions in Europe is part of the wider history of the lion species complex. The rediscovery and confirmation of their presence in Europe, already known by myths, historical accounts and ancient art, was made possible by the finds of fossils of Pleistocene, Holocene and Ancient lions excavated in Europe since the early 19th century. The first excavated lion fossil was found in southern Germany, and described by Georg August Goldfuss using the scientific name Felis spelaea. It probably dates to the Würm glaciation, and is 191,000 to 57,000 years old. Since then, older lion skull fragments were excavated in Germany and in other parts of Europe, including in Western Europe. Some of them were described by Wilhelm von Reichenau under Felis fossilis in 1906.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. One of the largest known elephant species, mature fully grown bulls on average had a shoulder height of 4 metres (13 ft) and a weight of 13 tonnes (29,000 lb). Straight-tusked elephants likely lived very similarly to modern elephants, with herds of adult females and juveniles and solitary adult males. The species was primarily associated with temperate and Mediterranean woodland and forest habitats, flourishing during interglacial periods, when its range would extend across Europe as far north as Great Britain and eastwards into Russia, while persisting in southern Europe during glacial periods. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Homo heidelbergensis and their Neanderthal successors.
Panthera fossilis is an extinct species of cat belonging to the genus Panthera, known from remains found in Eurasia spanning the Middle Pleistocene and possibly into the Early Pleistocene.
Cave hyena are extinct species or subspecies of hyena known from Eurasia, which ranged from Western Europe to eastern Asia and Siberia. It is one of the best known mammals of the Ice Age and is well represented in many European bone caves. It preyed on large mammals, and was responsible for the accumulation of hundreds of large Pleistocene mammal bones in areas including horizontal caves, sinkholes, mud pits, and muddy areas along rivers.
The Tremarctinae or short-faced bears is a subfamily of Ursidae that contains one living representative, the spectacled bear of South America, and several extinct species from four genera: the Florida spectacled bear, the North American giant short-faced bears Arctodus, the South American giant short-faced bear Arctotherium as well as Plionarctos(P. edensis and P. harroldorum), which is thought to be ancestral to the other three genera. Of these, the giant short-faced bears may have been the largest ever carnivorans in the Americas. The group is thought to have originated in eastern North America, and then invaded South America as part of the Great American Interchange. Most short-faced bears became extinct at the end of the Pleistocene.
Drachenhöhle or Drachenhöhle Mixnitz is a 542 m (1,778 ft) long cave with a 20 m (66 ft) wide and 12 m (39 ft) high entrance near Mixnitz, Styria, Austria, south-east of Bruck an der Mur located at an elevation of 950 m (3,120 ft) above sea level. Cave bear of the species and other bone fossils that people found during the Middle Ages were deemed to be the bones of dragons, a belief that culminated in the saga of the "Dragon slayer of Mixnitz". The cave is one of the largest caves in the Alps where bears occupied an area that stretched over a length of way over 500 m (1,600 ft), by an average width of up to 40 m (130 ft) and a height of 10 to 15 m.
Ursus deningeri is an extinct species of bear, endemic to Eurasia during the Pleistocene for approximately 1.7 million years, from 1.8 to 0.1 million years ago. The range of this bear has been found to encompass both Europe and Asia, demonstrating the ability of the species to adapt to many Pleistocene environments.
The Etruscan bear is an extinct species of bear, endemic to Europe, Asia and North Africa during the Early Pleistocene, living from approximately 2.2 million to around 1.4-1.2 million years ago.
Arctotherium is an extinct genus of the Pleistocene short-faced bears endemic to Central and South America. Arctotherium migrated from North America to South America during the Great American Interchange, following the formation of the Isthmus of Panama during the late Pliocene. The genus consists of one early giant form, A. angustidens, and several succeeding smaller species, which were within the size range of modern bears. Arctotherium was adapted to open and mixed habitat. They are genetically closer to the spectacled bear, than to Arctodus of North America, implying the two extinct forms evolved large size in a convergent manner.
Dusicyon avus is an extinct species of cerdocyonine canid in the genus Dusicyon, native to South America during the Pleistocene and Holocene epochs. It was medium to large, about the size of a German shepherd. It was closely related to the Falkland Islands wolf or warrah (Dusicyon australis), which descended from a population of D. avus. It appears to have survived until very recently, perhaps 400 years ago.
Ursus rossicus is an extinct species of bear that lived in the steppe regions of northern Eurasia and Siberia during the Pleistocene.
The small Sirgenstein Cave, German: Sirgensteinhöhle is situated 565 m (1,854 ft) above sea level inside the 20 m (66 ft) high Sirgenstein, a limestone rock. The cave sits 35 m (115 ft) above the Ach River valley bottom in the central Swabian Jura, southern Germany. Archaeologist R. R. Schmidt excavated the site in 1906 during which he identified indices of prehistoric human presence. He recorded the complete stratigraphic sequence of Palaeolithic and Neolithic origin. In his 1910 analysis Schmidt inspired future archaeologists with his pioneering concept of including the excavation site within its geographic region, contextualizing it within a wide scientific spectrum and demonstrated valuable results as he correlated the Sirgenstein layer structure to those of prehistoric sites in France. Mammoth ivory beads dating from 39,000 to 35,000 years ago have been uncovered at the cave. Because of its historical and cultural significance and its testimony to the development of Paleolithic art, the cave was inscribed on the UNESCO World Heritage List as part of the Caves and Ice Age Art in the Swabian Jura site in 2017.