Ursus rossicus Temporal range: Middle to Late Pleistocene, | |
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Skeleton in Zoological Museum, Saint Petersburg | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Ursidae |
Genus: | Ursus |
Species: | †U. rossicus |
Binomial name | |
†Ursus rossicus Borissiak, 1930 | |
Ursus rossicus (the Pleistocene small cave bear ) [1] is an extinct species of bear that lived in the steppe regions of northern Eurasia and Siberia during the Pleistocene.
Vereshchagin discovered the first U. rossicus remains in the Altai Mountains in 1973. Mandibles of the bear were found in Bachatsk Quarry, Krasni Yar (in Tomsk Province) and Mokhovo Quarry; skull fragments were found in Krasni Yar. [1] More fossil remains of U. rossicus were discovered in the famous Denisova Cave. [2]
The small cave bear had a very broad, domed skull with a steep forehead. Its stout body had long thighs, massive shins and in-turning feet, making it similar in skeletal structure to the brown bear. [3] Cave bears were comparable in size to the largest modern-day bears. [4]
U. rossicus dental specimens when separated into groups of different ontogenetic stages do not show significant variations in microwear, suggesting the diets of these bears stayed fairly constant over the course of their lifetimes. [5]
Cave bear teeth show greater wear than most modern bear species, suggesting a diet of tough materials. However, tubers and other gritty food, which cause distinctive tooth wear in modern brown bears, do not appear to have constituted a major part of cave bears' diets on the basis of dental microwear analysis. [6]
The morphological features of the cave bear chewing apparatus, including loss of premolars, have long been suggested to indicate their diets displayed a higher degree of herbivory than the Eurasian brown bear. Indeed, a solely vegetarian diet has been inferred on the basis of tooth morphology. Results obtained on the stable isotopes of cave bear bones also point to a largely vegetarian diet in having low levels of nitrogen-15 and carbon-13, [7] which are accumulated at a faster rate by carnivores as opposed to herbivores.
Cave bears of the last ice age lacked the usual two or three premolars present in other bears; to compensate, the last molar is very elongated, with supplementary cusps. [8]
The cave bear is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum.
The woolly rhinoceros is an extinct species of rhinoceros that inhabited northern Eurasia during the Pleistocene epoch. The woolly rhinoceros was a member of the Pleistocene megafauna. The woolly rhinoceros was covered with long, thick hair that allowed it to survive in the extremely cold, harsh mammoth steppe. It had a massive hump reaching from its shoulder and fed mainly on herbaceous plants that grew in the steppe. Mummified carcasses preserved in permafrost and many bone remains of woolly rhinoceroses have been found. Images of woolly rhinoceroses are found among cave paintings in Europe and Asia. The species range contracted towards Siberia beginning around 17,000 years ago, with the youngest known records being around 14,000 years old in northeast Siberia, coinciding with the Bølling–Allerød warming, which likely disrupted its habitat, with environmental DNA records possibly extending the range of the species around 9,800 years ago. Its closest living relative in the Sumatran rhinoceros.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.
Panthera spelaea, commonly known as the cave lion is an extinct Panthera species native to Eurasia and northwest North America during the Pleistocene epoch. Genetic analysis of ancient DNA has revealed that while closely related, it was a distinct species genetically isolated from the modern lion, with the genetic divergence between the two species estimated at around 500,000 years ago. The earliest fossils of the P. spelaea lineage in Eurasia date to around 700,000 years ago. It is closely related and probably ancestral to the American lion. The species ranged from Western Europe to eastern Beringia in North America, and was a prominent member of the mammoth steppe fauna, and an important apex predator across its range. It became extinct about 13,000 years ago. It closely resembled living lions with a coat of yellowish-grey fur, though unlike living lions males appear to have lacked manes.
The American lion is an extinct pantherine cat native to North America during the Late Pleistocene from around 130,000 to 12,800 years ago. Genetic evidence suggests that its closest living relative is the lion, with the American lion representing an offshoot from the lineage of the largely Eurasian cave lion, from which it is suggested to have split around 165,000 years ago. Its fossils have been found across North America, from Canada to Mexico. It was about 25% larger than the modern lion, making it one of the largest known felids to ever exist, and an important apex predator.
Gigantopithecus is an extinct genus of ape that lived in southern China from 2 million to approximately 300,000-200,000 years ago during the Early to Middle Pleistocene, represented by one species, Gigantopithecus blacki. Potential identifications have also been made in Thailand, Vietnam, and Indonesia. The first remains of Gigantopithecus, two third molar teeth, were identified in a drugstore by anthropologist Ralph von Koenigswald in 1935, who subsequently described the ape. In 1956, the first mandible and more than 1,000 teeth were found in Liucheng, and numerous more remains have since been found in at least 16 sites. Only teeth and four mandibles are known currently, and other skeletal elements were likely consumed by porcupines before they could fossilise. Gigantopithecus was once argued to be a hominin, a member of the human line, but it is now thought to be closely allied with orangutans, classified in the subfamily Ponginae.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. One of the largest known elephant species, mature bulls on average had a shoulder height of 3.81–4.2 metres (12.5–13.8 ft) and a weight of 10.8–15 tonnes (24,000–33,000 lb). Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.
Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.
Sinomastodon is an extinct gomphothere genus known from the Late Miocene to Early Pleistocene of Asia, including China, Japan, Thailand, Myanmar, Indonesia and probably Kashmir.
Archaeolemur is an extinct genus of subfossil lemurs known from the Holocene epoch of Madagascar. Archaeolemur is one of the most common and well-known of the extinct giant lemurs as hundreds of its bones have been discovered in fossil deposits across the island. It was larger than any extant lemur, with a body mass of approximately 18.2–26.5 kg (40–58 lb), and is commonly reconstructed as the most frugivorous and terrestrial of the fossil Malagasy primates. Colloquially known as a "monkey lemur," Archaeolemur has often been compared with anthropoids, specifically the cercopithecines, due to various morphological convergences. In fact, it was even misidentified as a monkey when remains were first discovered. Following human arrival to Madagascar just over 2000 years ago, many of the island’s megafauna went extinct, including the giant lemurs. Radiocarbon dating indicates that Archaeolemur survived on Madagascar until at least 1040-1290 AD, outliving most other subfossil lemurs.
Arctodus is an extinct genus of short-faced bear that inhabited North America during the Pleistocene. There are two recognized species: the lesser short-faced bear and the giant short-faced bear. Of these species, A. simus was larger, is known from more complete remains, and is considered one of the most charismatic of North America's megafauna. A. pristinus was largely restricted to the Early Pleistocene of the eastern United States, whereas A. simus had a broader range, with most finds being from the Late Pleistocene of the United States, Mexico and Canada. A. simus evolved from A. pristinus, but both species likely overlapped in the Middle Pleistocene. Both species are relatively rare in the fossil record.
Ursus deningeri is an extinct species of bear, endemic to Eurasia during the Pleistocene for approximately 1.7 million years, from 1.8 to 0.1 million years ago.
The narrow-nosed rhinoceros, also known as the steppe rhinoceros is an extinct species of rhinoceros belonging to the genus Stephanorhinus that lived in western Eurasia, including Europe, as well as North Africa during the Pleistocene. It first appeared in Europe around 500,000 years ago during the Middle Pleistocene and survived there until at least 34,000 years Before Present.
Stephanorhinus kirchbergensis, also known as Merck's rhinoceros or the forest rhinoceros, is an extinct species of rhinoceros belonging to the genus Stephanorhinus from the Middle to Late Pleistocene of Eurasia. Its range spanned from Western Europe to Eastern Asia. Among the last members of the genus, it co-existed alongside Stephanorhinus hemitoechus in the western part of its range.
The cave wolf is an extinct glacial mammoth steppe-adapted white wolf that lived during the Middle Pleistocene to the Late Pleistocene. It inhabited Europe, where its remains have been found in many caves. Its habitat included the mammoth steppe grasslands and boreal needle forests. This large wolf was short-legged compared to its body size, yet its leg size is comparable with that of the Arctic wolf C. l. arctos. Genetic evidence suggests that Late Pleistocene European wolves shared high genetic connectivity with contemporary wolves from Siberia, with continual gene flow from Siberian wolves into European wolves over the course of the Late Pleistocene. Modern European wolves derive most of their ancestry from Siberian wolf populations that expanded into Europe during and after the Last Glacial Maximum, but retain a minor fraction of their ancestry from earlier Late Pleistocene European wolves.
Ursus dolinensis is an extinct mammalian carnivore species of the Ursidae family. Its fossilized remains were unearthed from the lowest layers of the stratigraphic sequence at the archaeological and paleontological site of Gran Dolina, that is a part of the Atapuerca Mountains complex in the Burgos province, northern Spain. The species was described by Nuria Garcia and Juan Luis Arsuaga in a 2001 publication. Skeletal fossils, mainly cranial fragments were recovered from the sediment units TD 3 and in particular TD 4. Presence in these layers suggests a chronology in between 900,000 and 780,000 years ago, which falls into the Calabrian stage of the early Pleistocene.
Ursus ingressus is an extinct species of the family Ursidae that lived in Central Europe during the Late Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found.
This paleomammalogy list records new fossil mammal taxa that were described during the year 2018, as well as notes other significant paleomammalogy discoveries and events which occurred during that year.
This paleomammalogy list records new fossil mammal taxa that were described during the year 2019, as well as notes other significant paleomammalogy discoveries and events which occurred during that year.
Protarctos is an extinct genus of basal ursine bear that lived in North America and Eurasia during the Pliocene and into Early Pleistocene.