Eurasian brown bear | |
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An adult female in Slovenia | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Ursidae |
Genus: | Ursus |
Species: | |
Subspecies: | U. a. arctos |
Trinomial name | |
Ursus arctos arctos |
The Eurasian brown bear (Ursus arctos arctos) is one of the most common subspecies of the brown bear, [2] and is found in much of Eurasia. It is also called the European brown bear, common brown bear, common bear, and colloquially by many other names. The genetic diversity of present-day brown bears (Ursus arctos) has been extensively studied over the years and appears to be geographically structured into five main clades based upon analysis of the mtDNA. [3]
The Eurasian brown bear has brown fur, which ranges from yellowish-brown to dark brown, red-brown, and almost black in some cases; albinism has also been recorded. [4] The fur is dense to varying degrees and the hair can grow up to 10 cm (3.9 in) in length. The head normally is quite round and has relatively small rounded ears, a wide skull, and a mouth equipped with 42 teeth, including predatory teeth. It has a powerful bone structure and large paws equipped with claws that can grow up to 10 cm (3.9 in) in length. The weight varies depending on habitat and the time of the year. A full-grown male weighs on average between 350 and 500 kg and reaches a maximum weight of 650 kg and length of nearly 2.5 m (8.2 ft). Females typically range between 150 and 300 kg and reach a maximum weight of 450 kg . [5] They have a lifespan of 20 to 30 years in the wild. [6]
Eurasian brown bears were used in Ancient Rome for fighting in arenas. The strongest bears apparently came from Caledonia and Dalmatia. [7]
In antiquity, the Eurasian brown bear was largely carnivorous, with 80% of its diet consisting of animal matter. However, as its habitat increasingly diminished, the portion of meat in its diet decreased with it until by the late Middle Ages, meat consisted of only 40% of its dietary intake. Today, meat makes up little more than 10–15% of its diet. [7] Whenever possible, the brown bear will consume sheep. [8]
Unlike in North America, where an average of two people a year are killed by bears, Scandinavia only has records of three fatal bear attacks within the last century. [9] In late 2019, brown bears killed three men in Romania in just over a month. [10]
The oldest fossils are from the Choukoutien, China, and date back about 500,000 years. [11] It is known from mtDNA studies that during the Pleistocene ice age it was too cold for the brown bear to survive in Europe except in three places: Russia, Spain, and the Balkans. [12] However, a newer study found that brown bears were present in France and Belgium during the Last Glacial Maximum as well, indicating they were not as restricted to southern refugia as previously thought. [13]
Modern research [14] has made it possible to track the origin of the subspecies. The species to which it belongs developed more than 500,000 years ago, and researchers have found that the Eurasian brown bear separated about 850,000 years ago, with one branch based in Western Europe and the other branch in Russia, Eastern Europe and Asia. [12] Through the research of mitochondrial DNA (mtDNA), researchers have found that the European family has divided into two clades—one in the Iberian Peninsula and the Balkans, the other in Russia. [12] [15]
There is a population in Scandinavia that includes bears of the western and eastern lineages. [15] By analyzing the mtDNA of the southern population, researchers have found that they have probably come from populations in the Pyrenees in Southern France and Spain and the Cantabrian Mountains (Spain). Bears from these populations spread to southern Scandinavia after the last ice age. The northern bear populations originate in the Finnish/Russian population. Probably their ancestors survived the ice age in the ice-free areas west of the Ural Mountains, and thereafter spread to Northern Europe. [12]
Brown bears could once be found across most of Eurasia, compared to the more limited range today. General habitats included areas such as grassland, sparsely vegetated land, and wetlands.
Although included as of Least Concern on the 2006 IUCN Red List of Threatened Species (which refers to the global species, not to the Eurasian brown bear specifically), local populations, specifically those in the European Union, are becoming increasingly scarce. [16] As the IUCN itself adds: "Least Concern does not always mean that species are not at risk. There are declining species that are evaluated as Least Concern."
The brown bear has long been extinct in the British Isles (at least 1,500 years ago, possibly even 3,000 years ago), [17] [18] Denmark (about 6,500 years ago), [19] the Netherlands (about 1,000 years ago, although later singles rarely wandered from Germany), [20] Belgium and Luxembourg, with more recent extinctions in Germany (in the year 1835, although singles wandering from Italy were recorded in 2006 and 2019), [21] [22] Switzerland (in 1904, although a single was seen in 1923 and since 2005 there has been an increasing number of sightings of wanderers from Italy), [23] [24] and Portugal (in 1843, although a wanderer from Spain was recorded in 2019). [25]
Globally, the largest population is found east of the Ural mountain range, in the large Siberian forests; brown bears are also present in smaller numbers in parts of central Asia.
The largest brown bear population in Europe is in Russia, where it has now recovered from an all-time low caused by intensive hunting. Populations in Baltoscandia are similarly, albeit slowly, increasing. They include almost 3,000 bears in Sweden, 2,000 in Finland, 1,100 in Estonia [26] and around 100 in Norway.
Large populations can also be found in Romania (around 6,000), Slovakia (around 1,200), Bosnia and Herzegovina, Croatia (1,200), Slovenia (1,100), [27] North Macedonia, Bulgaria, Poland, Turkey, and Georgia.
Small but still significant populations can also be found in Albania, Greece, Serbia and Montenegro. [28] In 2005, there were an estimated 200 in Ukraine; these populations are part of two distinct metapopulations: the Carpathian with over 5000 individuals, and the Dinaric-Pindos (Balkans) with around 3000 individuals. [29]
There is a small but growing population (at least 70 bears) [30] in the Pyrenees, on the border between Spain and France, which was once on the edge of extinction, [31] as well as two subpopulations in the Cantabrian Mountains in Spain (amounting to around 250 individuals). [32] There are also populations totalling around 100 bears in the Abruzzo, South Tyrol and Trentino regions of Italy. [33] Bears from the aforementioned Italian regions occasionally cross over to bordering Switzerland, [34] [35] [36] which has not hosted a native population since its last bear was shot and killed in Graubünden in 1904. [37]
Outside Europe and Russia/the CIS, clades of brown bear persist in small, isolated, and for the most part highly threatened populations in Iran, [38] Afghanistan, Pakistan, parts of northwest India and central China, and on the island of Hokkaido in Japan. [39] [40]
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The historic distribution of bears and the impression the Eurasian brown bear has made on people are reflected in the names of several localities (some notable examples include Bern, Medvednica, Otepää and Ayu-Dag), as well as personal names—for example, Xiong, Bernard, Arthur, Ursula, Urs, Ursicinus, Orsolya, Björn, Nedved, Medvedev, and Otso.
Bears of this subspecies appear very frequently in the fairy tales and fables of Europe, in particular, tales collected by Jakob and Wilhelm Grimm. The European brown bear was once common in Germany and alpine lands like Northern Italy, Eastern France, and most of Switzerland, and thus appears in tales of various dialects of German.
The bear is traditionally regarded as the symbol of Russian (military and political) might. It is also Finland's national animal; [41] and in Croatia, a brown bear is depicted on the reverse of the Croatian 5 kuna coin, minted from 1993 to 2023. [42]
The brown bear is a large bear native to Eurasia and North America. Of the carnivorans, it is rivaled in size only by its closest relative, the polar bear, which is much less variable in size and slightly bigger on average. The brown bear is a sexually dimorphic species, as adult males are larger and more compactly built than females. The fur ranges in color from cream to reddish to dark brown. It has evolved large hump muscles, unique among bears, and paws up to 21 cm (8.3 in) wide and 36 cm (14 in) long, to effectively dig through dirt. Its teeth are similar to those of other bears and reflect its dietary plasticity.
The cave bear is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum.
The Eurasian lynx is one of the four extant species within the medium-sized wild cat genus Lynx. It is widely distributed from Northern, Central and Eastern Europe to Central Asia and Siberia, the Tibetan Plateau and the Himalayas. It inhabits temperate and boreal forests up to an elevation of 5,500 m (18,000 ft). Despite its wide distribution, it is threatened by habitat loss and fragmentation, poaching and depletion of prey.
Ursus is a genus in the family Ursidae (bears) that includes the widely distributed brown bear, the polar bear, the American black bear, and the Asian black bear. The name is derived from the Latin ursus, meaning bear.
The California grizzly bear, also known as the California golden bear, is an extinct population of the brown bear, generally known as the grizzly bear. "Grizzly" could have meant "grizzled" – that is, with golden and grey tips of the hair – or "fear-inspiring". Nonetheless, after careful study, naturalist George Ord formally classified it in 1815 – not for its hair, but for its character – as Ursus horribilis. Genetically, North American brown bears are closely related; in size and coloring, the California grizzly bear was much like the Kodiak bear of the southern coast of Alaska. The grizzly became a symbol of the Bear Flag Republic, a moniker that was attached to the short-lived attempt by a group of U.S. settlers to break away from Mexico in 1846. Later, this rebel flag became the basis for the state flag of California, and then California was known as the "Bear State".
The Atlas bear or North African bear was a population of brown bear native to North Africa that became extinct in historical times.
The Syrian brown bear is a medium-sized and endangered subspecies of Eurasian brown bear native to the Middle East and West-Central Asia, particularly around the Caucasus Mountains.
The Himalayan brown bear, also known as the Himalayan red bear or isabelline bear, is a subspecies of the brown bear occurring in the western Himalayas. It is the largest mammal in the region, males reaching up to 2.2 m long, while females are a little smaller. It is omnivorous and hibernates in dens during the winter.
The Gobi bear, known in Mongolian as the Mazaalai (Мазаалай), is a subspecies of the brown bear that is found in the Gobi Desert of Mongolia. It is listed as critically endangered by the Mongolian Redbook of Endangered Species and by IUCN standards. Currently, there are only 31 bears left in the Mongolian Gobi Desert; through long-term genetic monitoring it is known that the population is relatively stable, however, the sex ratio is highly skewed towards males. Gobi bears are separated by enough distance from other brown bear populations to achieve reproductive isolation. In 1959, hunting of the animal was prohibited in order to preserve the dying subspecies.
The Marsican brown bear, also known as the Apennine brown bear, and orso bruno marsicano in Italian, is a critically endangered population of the Eurasian brown bear, with a range restricted to the Parco Nazionale d'Abruzzo, Lazio e Molise, and the surrounding region in Italy. The Marsican brown bear differs slightly from other brown bears in its appearance and hibernation techniques. The bear's popular name is derived from Marsica, a historic area of the modern-day region of Abruzzo where the bear has long had a significant presence.
The Mexican grizzly bear is an extinct population of the grizzly bear in the Southwestern United States and Mexico.
Bears have been hunted since prehistoric times for their meat and fur. In addition to being a source of food, in modern times they have been favored by big game hunters due to their size and ferocity. Bear hunting has a vast history throughout Europe and North America, and hunting practices have varied based on location and type of bear.
The Cantabrian brown bear, Iberian brown bear, or Iberian bear is a population of Eurasian brown bears living in the Cantabrian Mountains of Spain.
The Ussuri brown bear, also known as the Ezo brown bear, Russian grizzly bear, or the black grizzly bear, is a subspecies of the brown bear or a population of the Eurasian brown bear. One of the largest brown bears, a very large Ussuri brown bear may approach the Kodiak bear in size. It is not to be confused with the North American grizzly bear.
Ursus deningeri is an extinct species of bear, endemic to Eurasia during the Pleistocene for approximately 1.7 million years, from 1.8 to 0.1 million years ago. The range of this bear has been found to encompass both Europe and Asia, demonstrating the ability of the species to adapt to many Pleistocene environments.
The Etruscan bear is an extinct species of bear, endemic to Europe, Asia and North Africa during the Early Pleistocene, living from approximately 2.2 million to around 1.4-1.2 million years ago.
The grizzly bear, also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America.
Formerly or currently considered subspecies or populations of brown bears have been listed as follows:
The brown bear is one of the most omnivorous animals in the world and has been recorded to consume the greatest variety of foods of any bear. Throughout life, this species is regularly curious about the potential of eating virtually any organism or object that they encounter. Certainly no other animal in their given ecosystems, short perhaps of other bear species and humans, can claim to feed on as broad a range of dietary opportunities. Food that is both abundant and easily obtained is preferred. Their jaw structure has evolved to fit their dietary habits. Their diet varies enormously throughout their differing areas based on opportunity. In spring, winter-provided carrion, grasses, shoots, sedges and forbs are the dietary mainstays for brown bears from almost every part of their distribution. Fruits, including berries, become increasingly important during summer and early autumn. Roots and bulbs become critical in autumn for some inland bear populations if fruit crops are poor. The dietary variability is illustrated in the western United States, as meat made up 51% of the average year-around diet for grizzly bears from Yellowstone National Park, while it made up only 11% of the year-around diet for grizzlies from Glacier National Park a few hundred miles to the north.
Brown bears were once native to Europe, much of Asia, the Atlas Mountains of Africa, and North America, but are now extirpated in some areas, and their populations have greatly decreased in other areas. There are approximately 200,000 brown bears left in the world. The largest population is in Russia, with 120,000 individuals. The brown bear occupies the largest range of habitats of any Ursus species with recorded observations in every temperate northern forest and at elevations as high as 5,000 m.