The brown bear is a large bear native to Eurasia and North America. Of the carnivorans, it is rivaled in size only by its closest relative, the polar bear, which is much less variable in size and slightly bigger on average. The brown bear is a sexually dimorphic species, as adult males are larger and more compactly built than females. The fur ranges in color from cream to reddish to dark brown. It has evolved large hump muscles, unique among bears, and paws up to 21 cm (8.3 in) wide and 36 cm (14 in) long, to effectively dig through dirt. Its teeth are similar to those of other bears and reflect its dietary plasticity.
The cave bear is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum.
The steppe bison or steppe wisent is an extinct species of bison. It was widely distributed across the mammoth steppe, ranging from Western Europe to eastern Beringia in North America during the Late Pleistocene. It is ancestral to all North American bison, including ultimately modern American bison. Three chronological subspecies, Bison priscus priscus, Bison priscus mediator, and Bison priscus gigas, have been suggested.
Panthera spelaea, commonly known as the cave lion, is an extinct Panthera species that was native to Eurasia and northwest North America during the Pleistocene epoch. Genetic analysis of ancient DNA has revealed that while closely related, it was a distinct species genetically isolated from the modern lion, with the genetic divergence between the two species estimated at around 500,000 years ago. The earliest fossils of the P. spelaea lineage in Eurasia date to around 700,000 years ago. It is closely related and probably ancestral to the American lion. The species ranged from Western Europe to eastern Beringia in North America, and was a prominent member of the mammoth steppe fauna, and an important apex predator across its range. It became extinct about 13,000 years ago. It closely resembled living lions with a coat of yellowish-grey fur though unlike extant lions, males appear to have lacked manes.
The American lion, with the species name meaning "savage" or "cruel", also called the North American lion) is an extinct pantherine cat native to North America during the Late Pleistocene from around 130,000 to 12,800 years ago. Genetic evidence suggests that its closest living relative is the lion, with the American lion representing an offshoot from the lineage of the largely Eurasian cave lion, from which it is suggested to have split around 165,000 years ago. Its fossils have been found across North America, from Canada to Mexico. It was about 25% larger than the modern lion, making it one of the largest known felids to ever exist, and an important apex predator.
The Eurasian brown bear is one of the most common subspecies of the brown bear, and is found in much of Eurasia. It is also called the European brown bear, common brown bear, common bear, and colloquially by many other names. The genetic diversity of present-day brown bears has been extensively studied over the years and appears to be geographically structured into five main clades based upon analysis of the mtDNA.
The Gobi bear, known in Mongolian as the Mazaalai (Мазаалай), is a subspecies of the brown bear that is found in the Gobi Desert of Mongolia. It is listed as critically endangered by the Mongolian Redbook of Endangered Species and by IUCN standards. Currently, there are only 31 bears left in the Mongolian Gobi Desert; through long-term genetic monitoring it is known that the population is relatively stable, however, the sex ratio is highly skewed towards males. Gobi bears are separated by enough distance from other brown bear populations to achieve reproductive isolation. In 1959, hunting of the animal was prohibited in order to preserve the dying subspecies.
The Cantabrian brown bear, Iberian brown bear, or Iberian bear is a population of Eurasian brown bears living in the Cantabrian Mountains of Spain.
Arctodus is an extinct genus of short-faced bear that inhabited North America during the Pleistocene. There are two recognized species: the lesser short-faced bear and the giant short-faced bear. Of these species, A. simus was larger, is known from more complete remains, and is considered one of the best known members of North America's extinct Ice Age megafauna. A. pristinus was largely restricted to the Early Pleistocene of the eastern United States, whereas A. simus had a broader range, with most finds being from the Late Pleistocene of the United States, Mexico and Canada. A. simus evolved from A. pristinus, but both species likely overlapped in the Middle Pleistocene. Both species are relatively rare in the fossil record.
Ursus maritimus tyrannus is a controversial extinct subspecies of polar bear.
The Etruscan bear is an extinct species of bear, endemic to Europe, Asia and North Africa during the Early Pleistocene, living from approximately 2.2 million to around 1.4-1.2 million years ago.
The Alaska Peninsula brown bear or "peninsular grizzly" is a colloquial nomenclature for a possible brown bear subspecies that lives in the coastal regions of southern Alaska. It may be a population of the mainland grizzly bear subspecies.
The Beringian wolf is an extinct population of wolf that lived during the Ice Age. It inhabited what is now modern-day Alaska, Yukon, and northern British Columbia. Some of these wolves survived well into the Holocene. The Beringian wolf is an ecomorph of the gray wolf and has been comprehensively studied using a range of scientific techniques, yielding new information on their prey species and feeding behaviors. It has been determined that these wolves are morphologically distinct from modern North American wolves and genetically basal to most modern and extinct wolves. The Beringian wolf has not been assigned a subspecies classification and its relationship with the extinct European cave wolf is not clear.
The ABC Islands bear or Sitka brown bear is a subspecies or population of brown bear that resides in Southeast Alaska and is found on Admiralty Island, Baranof Island, and Chichagof Island in Alaska, and a part of the Alexander Archipelago. It has a unique genetic structure that relates them not only to brown bears, but also to polar bears. Its habitat exists within the Tongass National Forest, which is part of the perhumid rainforest zone.
The cave wolf is an extinct glacial mammoth steppe-adapted wolf that lived during the Middle Pleistocene to the Late Pleistocene. It inhabited Europe, where its remains have been found in many caves. Its habitat included the mammoth steppe grasslands and boreal needle forests. This large wolf was short-legged compared to its body size, yet its leg size is comparable with that of the Arctic wolf C. l. arctos. Genetic evidence suggests that Late Pleistocene European wolves shared high genetic connectivity with contemporary wolves from Siberia, with continual gene flow from Siberian wolves into European wolves over the course of the Late Pleistocene. Modern European wolves derive most of their ancestry from Siberian wolf populations that expanded into Europe during and after the Last Glacial Maximum, but retain a minor fraction of their ancestry from earlier Late Pleistocene European wolves.
Ursus dolinensis is an extinct mammalian carnivore species of the Ursidae family. Its fossilized remains were unearthed from the lowest layers of the stratigraphic sequence at the archaeological and paleontological site of Gran Dolina, that is a part of the Atapuerca Mountains complex in the Burgos province, northern Spain. The species was described by Nuria Garcia and Juan Luis Arsuaga in a 2001 publication. Skeletal fossils, mainly cranial fragments were recovered from the sediment units TD 3 and in particular TD 4. Presence in these layers suggests a chronology in between 900,000 and 780,000 years ago, which falls into the Calabrian stage of the early Pleistocene.
Ursus ingressus is an extinct species of the family Ursidae that lived in Central Europe during the Late Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found.
Formerly or currently considered subspecies or populations of brown bears have been listed as follows:
Protarctos is an extinct genus of basal ursine bear that lived in North America and Eurasia during the Pliocene and into Early Pleistocene.
