| Etruscan bear Temporal range: | |
|---|---|
 | |
| Fossils | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Carnivora |
| Family: | Ursidae |
| Genus: | Ursus |
| Species: | †U. etruscus |
| Binomial name | |
| †Ursus etruscus Cuvier, 1823 | |
The Etruscan bear (Ursus etruscus) is an extinct species of bear, endemic to Europe, Asia and North Africa during the Early Pleistocene, living from approximately 2.2 million to around 1.4-1.2 million years ago. [1]
The Etruscan bear appears to have evolved from Ursus minimus and gave rise to the modern brown bear, Ursus arctos , and the extinct cave bear, Ursus spelaeus . [2] The range of Etruscan bears was mostly limited to continental Europe, with specimens also recovered in the Great Steppe region of Eurasia. Fossil evidence for the Etruscan bear was recovered in Palestine, Greece, [3] Croatia, and Tuscany, Italy.
Some scientists have proposed that the early, small variety of U. etruscus of the middle Villafranchian era survives in the form of the modern Asian black bear. [4]
Not unlike the brown bears of Europe in size, it had a full complement of premolars, a trait carried from the genus Ursavus .
Ursus etruscus, like modern brown bears, is suggested to have been omnivorous, [5] with the diet of some populations suggested to heavily include fish. [6]
Sites and specimen ages:
The brown bear is a large bear native to Eurasia and North America. Of the carnivorans, it is rivaled in size only by its closest relative, the polar bear, which is much less variable in size and slightly bigger on average. The brown bear is a sexually dimorphic species, as adult males are larger and more compactly built than females. The fur ranges in color from cream to reddish to dark brown. It has evolved large hump muscles, unique among bears, and paws up to 21 cm (8.3 in) wide and 36 cm (14 in) long, to effectively dig through dirt. Its teeth are similar to those of other bears and reflect its dietary plasticity.
Dinofelis is an extinct genus of machairodontine, usually classified in the tribe Metailurini. It was widespread in Europe, Asia, Africa and North America from 5 million to about 1.2 million years ago. Fossils very similar to Dinofelis from Lothagam range back to around 8 million years ago, in the Late Miocene.
The cave bear is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum.
Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998), and Jin et al. (2007).
Pachycrocuta is an extinct genus of prehistoric hyenas. The largest and most well-researched species is Pachycrocuta brevirostris, colloquially known as the giant short-faced hyena as it stood about 90–100 cm (35–39 in) at the shoulder and it is estimated to have averaged 110 kg (240 lb) in weight, approaching the size of a lioness, making it the largest known hyena. Pachycrocuta first appeared during the late Miocene. By 800,000 years ago, it became locally extinct in Europe, with it surviving in East Asia until at least 500,000 years ago, and possibly later elsewhere in Asia.
Puma pardoides is an extinct prehistoric cat in the genus Puma known from fossils found across Eurasia.
Xenocyon is an extinct group of canids, either considered a distinct genus or a subgenus of Canis. The group includes Canis (Xenocyon) africanus, Canis (Xenocyon) antonii and Canis (Xenocyon) falconeri that gave rise to Canis (Xenocyon) lycanoides. The hypercarnivorous Xenocyon is thought to be closely related and possibly ancestral to modern dhole and the African wild dog, as well as the insular Sardinian dhole.
Leptobos is an extinct genus of large bovine, known from the Late Pliocene and Early Pleistocene of Eurasia, extending from the Iberian Peninsula to northern China. Species of Leptobos weighed on average 320 kilograms (710 lb). The dietary preference across the genus includes species that were browsers, grazers and mixed feeders. The first appearance of Leptobos in Europe around 3.6-3.5 million years ago is considered to define the beginning of the Villafranchian European faunal stage. Leptobos is considered to be closely related to the insular genus Epileptobos from the Pleistocene of Java, and is considered to be ancestral to Bison. Leptobos became extinct after being replaced by their descendant Bison during the Early Pleistocene, after a period of temporal overlap. "Leptobos" syrticus from Libya likely belongs in a different genus.
Ursus deningeri is an extinct species of bear, endemic to Eurasia during the Pleistocene for approximately 1.7 million years, from 1.8 to 0.1 million years ago. The range of this bear has been found to encompass both Europe and Asia, demonstrating the ability of the species to adapt to many Pleistocene environments.
Ursus minimus is an extinct species of bear, endemic to Europe during the Pliocene and Pleistocene, living from 5.3 to 1.8 Mya, existing for about 3.5 million years.
Stephanorhinus is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Late Pliocene to Late Pleistocene. Species of Stephanorhinus were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. The last two species of Stephanorhinus – Merck's rhinoceros and the narrow-nosed rhinoceros – went extinct during the last glacial period.
Pliocrocuta is an extinct genus of hyena. It contains the species Pliocrocuta perrieri, known from the Pliocene to Early Pleistocene of Eurasia and possibly Africa. It is possibly ancestral to Pachycrocuta. The species is estimated to have weighed around 56 kilograms (123 lb) on average, with its skull showing evidence for adaptation to bone cracking.
The cave wolf is an extinct glacial mammoth steppe-adapted wolf that lived during the Middle Pleistocene to the Late Pleistocene. It inhabited Europe, where its remains have been found in many caves. Its habitat included the mammoth steppe grasslands and boreal needle forests. This large wolf was short-legged compared to its body size, yet its leg size is comparable with that of the Arctic wolf C. l. arctos. Genetic evidence suggests that Late Pleistocene European wolves shared high genetic connectivity with contemporary wolves from Siberia, with continual gene flow from Siberian wolves into European wolves over the course of the Late Pleistocene. Modern European wolves derive most of their ancestry from Siberian wolf populations that expanded into Europe during and after the Last Glacial Maximum, but retain a minor fraction of their ancestry from earlier Late Pleistocene European wolves.
Ursus dolinensis is an extinct mammalian carnivore species of the Ursidae family. Its fossilized remains were unearthed from the lowest layers of the stratigraphic sequence at the archaeological and paleontological site of Gran Dolina, that is a part of the Atapuerca Mountains complex in the Burgos province, northern Spain. The species was described by Nuria Garcia and Juan Luis Arsuaga in a 2001 publication. Skeletal fossils, mainly cranial fragments were recovered from the sediment units TD 3 and in particular TD 4. Presence in these layers suggests a chronology in between 900,000 and 780,000 years ago, which falls into the Calabrian stage of the early Pleistocene.
The cave lynx or Mediterranean cave lynx is an extinct felid species that lived during the Pleistocene. It is controversially discussed to be a subspecies of the modern Iberian lynx and to be a junior synonym of this species.
Ursus ingressus is an extinct species of the family Ursidae that lived in Central Europe during the Late Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found.
Canis arnensis, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. Canis arnensis has been described as a small jackal-like canid. Its anatomy and morphology relate it more to the modern golden jackal than to the larger Etruscan wolf of that time. It is probably the ancestor of modern jackals.
Canis etruscus, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. The Etruscan wolf is described as a small wolf-like dog. It is widely agreed to be the ancestor of Canis mosbachensis, and thus ultimately the modern grey wolf.
Canis mosbachensis is an extinct wolf that inhabited Europe from the late Early Pleistocene to the Middle Pleistocene, around 1.4 million to 400,000 years ago. Canis mosbachensis is widely considered to have descended from the earlier Canis etruscus, and to be the ancestor of the living grey wolf with some considering it as a subspecies of the wolf as Canis lupus mosbachensis. The morphological distinction between C. mosbachensis and C. lupus has historically been vague, and attribution of fossils to C. mosbachensis or to C. lupus around the transition time between the two species is ambiguous.
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