Agriotheriini

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Agriotheriini
Temporal range: Tortonian–Gelasian
Indarctos arctoides - Batallones 3 fossil site, Torrejon de Velasco, Madrid, Spain.jpg
Skeletal remains of Indarctos arctoides displayed in National Archaeological Museum (Madrid)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Ursidae
Subfamily: Ailuropodinae
Tribe: Agriotheriini
Hendey, 1972
Genera
Synonyms
  • AgriotheriinaeKretzoi, 1929
  • IndarctiniAbella et al., 2012

Agriotheriini is an extinct tribe of ailuropodine bears from the middle Neogene to early Quaternary periods, with fossils found from Eurasia, Africa, and North America.

Taxonomy

The tribe Agriotheriini consists of the three (perhaps four) genera

The taxonomy of these bears has variously placed some of the genera in other bear lineages such as Hemicyoninae and Ursavinae. [1] [2]

Recent papers support their inclusion with giant pandas as members of Ailuropodinae based on diagnostic features like

They are unlike their closest living relative, the giant panda (Ailuropoda melanoleuca) which evolved into a highly specialized bamboo-eater: The evolution of agriotheriins lead to the group becoming large, hypercarnivorous bears that had adaptations of cursoriality unique in the evolutionary history of bears. [3] [4]

Related Research Articles

<span class="mw-page-title-main">Bear</span> Family of carnivoran mammals

Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout most of the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, and Eurasia. Common characteristics of modern bears include large bodies with stocky legs, long snouts, small rounded ears, shaggy hair, plantigrade paws with five nonretractile claws, and short tails.

<span class="mw-page-title-main">Carnivora</span> Order of mammals

Carnivora is an order of placental mammals that have specialized in primarily eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the fifth largest order of mammals, comprising at least 279 species on every major landmass and in a variety of habitats, ranging from the cold Polar regions of Earth the hyper-arid region of the Sahara Desert to the open seas. They have a very large array of different body plans with a wide diversity of shapes and sizes.

<span class="mw-page-title-main">Canidae</span> Family of mammals

Canidae is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid. The family includes three subfamilies: the Caninae, the extinct Borophaginae and Hesperocyoninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other species.

<span class="mw-page-title-main">Ursinae</span> Subfamily of bears

Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998), and Jin et al. (2007).

<span class="mw-page-title-main">Amphicyonidae</span> Extinct family of carnivores

Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene, spread to Europe by the late Eocene, and further spread to Asia and Africa by the early Miocene. They had largely disappeared worldwide by the late Miocene, with the latest recorded species at the end of the Miocene in Africa. They were among the first carnivorans to evolve large body size. Amphicyonids are colloquially referred to as "bear-dogs".

<span class="mw-page-title-main">Caniformia</span> Suborder of mammals

Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs, bears, raccoons, and mustelids. The Pinnipedia are also assigned to this group. The center of diversification for the Caniformia is North America and northern Eurasia. Caniformia stands in contrast to the other suborder of Carnivora, the Feliformia, the center of diversification of which was in Africa and southern Asia.

<span class="mw-page-title-main">Ailuridae</span> Family of carnivores

Ailuridae is a family in the mammal order Carnivora. The family consists of the red panda and its extinct relatives.

<span class="mw-page-title-main">Creodonta</span> Former order of extinct flesh-eating placental mammals

Creodonta is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ancestral to the modern Carnivora, this order is now usually considered a polyphyletic assemblage of two different groups, the Oxyaenids and the Hyenodonts, not a natural group. Oxyaenids are first known from the Palaeocene of North America, while hyaenodonts hail from the Palaeocene of Africa.

<span class="mw-page-title-main">Ailuropodinae</span> Subfamily of bears

Ailuropodinae is a subfamily of Ursidae that contains only one extant species, the giant panda of China. The fossil record of this group has shown that various species of pandas were more widespread across the Holarctic, with species found in places such as Europe, much of Asia, North America and even Africa. The earliest pandas were not unlike other modern bear species in that they had an omnivorous diet but by around 2.4 million years ago, pandas have evolved to be more herbivorous.

<span class="mw-page-title-main">Tremarctinae</span> Subfamily of bears

The Tremarctinae or short-faced bears is a subfamily of Ursidae that contains one living representative, the spectacled bear of South America, and several extinct species from four genera: the Florida spectacled bear, the North American giant short-faced bears Arctodus, the South American giant short-faced bear Arctotherium as well as Plionarctos(P. edensis and P. harroldorum), which is thought to be ancestral to the other three genera. Of these, the giant short-faced bears may have been the largest ever carnivorans in the Americas. The group is thought to have originated in eastern North America, and then invaded South America as part of the Great American Interchange. Most short-faced bears became extinct at the end of the Pleistocene.

<i>Simocyon</i> Extinct genus of carnivores

Simocyon is a genus of extinct carnivoran mammal in the family Ailuridae. Simocyon, which was about the size of a mountain lion, lived in the late Miocene and early Pliocene epochs, and has been found in Europe, Asia, and rarely, North America and Africa.

Parictis is an extinct arctoid belonging to the family Subparictidae.

<i>Dissacus</i> Extinct family of mammals

Dissacus is a genus of extinct carnivorous jackal to coyote-sized mammals within the family Mesonychidae, an early group of hoofed mammals that evolved into hunters and omnivores. Their fossils are found in Paleocene to Early Eocene aged strata in France, Asia and southwest North America, from 66 to 50.3 mya, existing for approximately 15.7 million years.

<span class="mw-page-title-main">Arctocyonidae</span> Extinct family of mammals

Arctocyonidae is as an extinct family of unspecialized, primitive mammals with more than 20 genera. Animals assigned to this family were most abundant during the Paleocene, but extant from the late Cretaceous to the early Eocene . Like most early mammals, their actual relationships are very difficult to resolve. No Paleocene fossil has been unambiguously assigned to any living order of placental mammals, and many genera resemble each other: generalized robust, not very agile animals with long tails and all-purpose chewing teeth, living in warm closed-canopy forests with many niches left vacant by the K-T extinction.

<i>Kolponomos</i> Extinct genus of mammals

Kolponomos is an extinct genus of carnivoran mammal that existed in the Late Arikareean North American Land Mammal Age, early Miocene epoch, about 20 million years ago. It was likely a marine mammal. The genus was erected in 1960 by Ruben A. Stirton, a paleontologist at the University of California Museum of Paleontology, Berkeley, for the species K. clallamensis, on the basis of a partial skull and jaw found on the Olympic Peninsula. At the time, Stirton questionably assigned it to Procyonidae, its systematic position remained problematic until the discovery of more fossils including a nearly complete cranium from the original locality of K. clallamensis which helped identify it as part of the group from which pinnipeds evolved.

<span class="mw-page-title-main">Giant tapir</span> Extinct species of mammal

The giant tapir is an extinct species of tapir that lived in southern China, Vietnam and Laos, with reports suggesting it also lived in Taiwan, Java, and potentially Borneo. The species has been recorded from Middle and Late Pleistocene. There is only weak evidence for a Holocene survival. Tapirus augustus was larger than any living tapir, with an estimated weight of about 623 kilograms (1,373 lb). The species was also placed in its own genus of Megatapirus, however, it is now conventionally placed within Tapirus.

<span class="mw-page-title-main">Hemicyoninae</span> Extinct subfamily of bears

Hemicyoninae is an extinct subfamily of Ursidae, often called dog bears. They were bear-like carnivorans living in Europe, North America, Africa and Asia during the Oligocene through Miocene epochs 33.9–5.3 Ma, existing for approximately 28.6 million years. They are sometimes classified as a separate family.

<span class="mw-page-title-main">Ursavini</span> Extinct tribe of bears

Ursavini is an extinct tribe of mammals of the family Ursidae (bears) endemic to North America, Europe, Africa, and Asia during the Miocene through the Pliocene, living from about 23—2.5 Mya, existing for roughly 20.5 million years.

<i>Arctotherium</i> Extinct genus of bears

Arctotherium is an extinct genus of the Pleistocene short-faced bears endemic to Central and South America. Arctotherium migrated from North America to South America during the Great American Interchange, following the formation of the Isthmus of Panama during the late Pliocene. The genus consists of one early giant form, A. angustidens, and several succeeding smaller species, which were within the size range of modern bears. Arctotherium was adapted to open and mixed habitat. They are genetically closer to the spectacled bear, than to Arctodus of North America, implying the two extinct forms evolved large size in a convergent manner.

Huracan is an extinct genus of agriotheriin ailuropodine from the Neogene period. Fossils were found from North America and Eastern Asia. Unlike its modern relative the giant panda, which is a bamboo specialist, Huracan was a hypercarnivorous genus of bear that had adaptations for cursoriality.

References

  1. Hunt, R.M. (1998). "Ursidae". In Jacobs, Louis; Janis, Christine M.; Scott, Kathleen L. (eds.). Evolution of Tertiary Mammals of North America. Vol. 1, Terrestrial carnivores, ungulates, and ungulate-like mammals. Cambridge, UK: Cambridge University Press. pp. 174–195. ISBN   0-521-35519-2.
  2. Jin, C.; Ciochon, R.L.; Dong, W.; Hunt, R.M., Jr.; Liu, J.; Jaeger, M.; Zhu, Q. (2007). "The first skull of the earliest giant panda". Proceedings of the National Academy of Sciences . 104 (26): 10932–10937. doi: 10.1073/pnas.0704198104 . PMC   1904166 . PMID   17578912.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  3. de Bonis, Louis; Merceron, Juan; Abella, Gildas; Begun, David (October 2017). "A new late Miocene ailuropodine (Giant Panda) from Rudabánya (North-central Hungary)". Geobios . 50 (5–6): 413–421. Bibcode:2017Geobi..50..413D. doi:10.1016/j.geobios.2017.09.003.
  4. Jiangzuo, Q.; Flynn, J.J.; Wang, S.; Hou, S.; Deng, T. (2023). "New fossil giant panda relatives (Ailuropodinae, Ursidae): A basal lineage of gigantic Mio-Pliocene cursorial carnivores". American Museum Novitates (3996): 1–71. doi: 10.1206/3996.1 . hdl:2246/7315. S2CID   257508340.