Zavacephale (meaning "origin head") is an extinct genus of pachycephalosaurian dinosaurs known from the Early CretaceousKhuren Dukh Formation of Mongolia. The genus contains a single species, Zavacephale rinpoche, known from a partial articulated skeleton including a nearly complete skull. It is both the oldest definitive pachycephalosaur known, as well as the most complete. Its skull bears a well-developed dome structure, as seen in most members of the clade.
The Zavacephaleholotype specimen, MPC-D 100/1209, was discovered in 2019 outcrops of the Khuren Dukh Formation in the Eastern Gobi Basin in Dundgobi Province, Mongolia. It was subsequently accessioned at the Institute of Paleontology (Mongolian Academy of Sciences) where it was prepared over the following years. The skeleton, which was found in partial articulation, is 54% complete based on element count, making it the most complete pachycephalosaur described. The specimen includes regions of the body not scientifically described in any other pachycephalosaur, including bones of the hand, a gastrolith mass, and a complete tail covered in ossified tendons . The skeleton also includes a complete skull and mandible, a scapulocoracoid , dorsal vertebrae , the sacrum , and most of the pelvic girdle and hindlimbs.[1] The fossil material was reported in conferenceabstracts in 2023 before its formal description.[2][3]
In September 2025, Tsogtbaatar Chinzorig and colleagues describedZavacephale rinpoche as a new genus and species of pachycephalosaurs based on these fossil remains. The generic name, Zavacephale, combines the Tibetan word zava meaning "root" or "origin" with the Greekkephale, meaning "head", referencing the basal position of the taxon within Pachycephalosauria. The specific name, rinpoche, is a Tibetan word meaning "precious one", alluding to the fact that the domed skull was found exposed in the rock outcrop, reminiscent of a cabochon gemstone.[1]
Description
Size compared to a human
The Zavacephale holotype belongs to an immature individual that was actively growing when it died, meaning it had not yet reached its full body size. This was determined based on the lack of fusion at the neurocentral sutures and between the scapula and coracoid, in addition to the lack of an external fundamental system and presence of limited remodelling of the tibia cortex. The skeleton is about 1 metre (3.3ft) long, and the animal likely weighed around 5.85 kilograms (12.9lb) in life.[1]
To test the relationships and affinities of Zavacephale, Chinzorig et al. (2025) compiled a new phylogenetic matrix derived from several past analyses. They recovered Zavacephale as a very basal member of the Pachycephalosauria, diverging after Wannanosaurus. As such, it is the sister taxon to all other pachycephalosaurs. These results are displayed in the cladogram below,[1] with node names following Madzia et al. (2021):[4]
Zavacephale is known from the upper portion of the lower Khuren Dukh Formation, which dates to around the Aptian to Albian ages of the early Cretaceous period. As such, it predates all other known pachycephalosaurs, which are restricted to the Late Cretaceous.[1] Other dinosaurs named from the formation include the hadrosauroid ornithopods Altirhinus and Choyrodon and the ornithomimosaur theropod Harpymimus. Choristoderes, turtles, and fish have also been described from the formation.[5][6]
References
1 2 3 4 5 Chinzorig, Tsogtbaatar; Takasaki, Ryuji; Yoshida, Junki; Tucker, Ryan T.; Buyantegsh, Batsaikhan; Mainbayar, Buuvei; Tsogtbaatar, Khishigjav; Zanno, Lindsay E. (2025-09-17). "A domed pachycephalosaur from the early Cretaceous of Mongolia". Nature: 1–8. doi:10.1038/s41586-025-09213-6. ISSN1476-4687.
↑ Chinzorig, Tsogtbaatar; Takasaki, Ryuji; Yoshida, Junki; Buyantegsh, Batsaikhan; Mainbayar, Buuvei; Zanno, Lindsay E. (2023). The oldest pachycephalosaur (Ornithischia: Marginocephalia) from the Early Cretaceous Khukhteeg Formation at Khuren Dukh locality of Mongolia. 14th Symposium on Mesozoic Terrestrial Ecosystems and Biota. Vol.306. pp.247–249. doi:10.1002/ar.25219.
↑ Matsumoto, Ryoko; Suzuki, Shigeru; Tsogtbaatar, Khisigjav; Evans, Susan E. (2008-11-26). "New material of the enigmatic reptile Khurendukhosaurus (Diapsida: Choristodera) from Mongolia". Naturwissenschaften. 96 (2): 233–242. doi:10.1007/s00114-008-0469-6. ISSN0028-1042.
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