2025 in archosaur paleontology

Last updated
List of years in archosaur paleontology
In paleontology
2022
2023
2024
2025
2026
2027
2028
In paleobotany
2022
2023
2024
2025
2026
2027
2028
In arthropod paleontology
2022
2023
2024
2025
2026
2027
2028
In paleoentomology
2022
2023
2024
2025
2026
2027
2028
In paleomalacology
2022
2023
2024
2025
2026
2027
2028
In reptile paleontology
2022
2023
2024
2025
2026
2027
2028
In paleomammalogy
2022
2023
2024
2025
2026
2027
2028
In paleoichthyology
2022
2023
2024
2025
2026
2027
2028

This article records new taxa of fossil archosaurs of every kind that are scheduled described during the year 2025, as well as other significant discoveries and events related to paleontology of archosaurs that are scheduled to occur in the year 2025.

Contents

Pseudosuchians

New pseudosuchian taxa

NameNoveltyStatusAuthorsAgeType localityLocationNotesImage

Kuttysuchus [1]

Gen. et sp. nov

Haldar, Ray & Bandyopadhyay

Late Triassic

Lower Dharmaram Formation

Flag of India.svg  India

An aetosaur belonging to the tribe Paratypothoracini. The type species is K. minori.

Pattisaura [2]

Gen. et sp. nov

Valid

Wu et al.

Late Triassic

Cooper Canyon Formation

Flag of the United States.svg  United States
(Flag of Texas.svg  Texas)

An early member of Crocodylomorpha. The type species is P. gracilis.

Pattisaura gracilis.jpg
Tewkensuchus [3] Gen. et sp. novBravo et al.Early Paleocene Salamanca Formation Flag of Argentina.svg  Argentina A sebecosuchian. The type species is T. salamanquensis

Thilastikosuchus [4]

Gen. et sp. nov

Valid

Carvalho et al.

Early Cretaceous

Quiricó Formation

Flag of Brazil.svg  Brazil

A notosuchian. The type species is T. scutorectangularis.

General pseudosuchian research

Aetosaur research

Crocodylomorph research

Non-avian dinosaurs

New dinosaur taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Ahvaytum [19]

Gen. et sp. nov

Lovelace et al.

Late Triassic (Carnian)

Popo Agie Formation

Flag of the United States.svg  United States
(Flag of Wyoming.svg  Wyoming)

An early saurischian, possibly a basal sauropodomorph. The type species is A. bahndooiveche.

Ahvaytum bahndooiveche.png

Archaeocursor [20]

Gen. et sp. nov

Valid

Yao et al.

Early Jurassic (SinemurianPliensbachian)

Ziliujing Formation

Flag of the People's Republic of China.svg  China

A basal ornithischian. The type species is A. asiaticus. Announced in 2024; the final article version was published in 2025.

Archaeocursor asiaticus.png

Chadititan [21]

Gen. et sp. nov

Valid

Agnolín et al.

Late Cretaceous (Campanian)

Anacleto Formation

Flag of Argentina.svg  Argentina

A rinconsaurian titanosaur. The type species is C. calvoi.

Chadititan calvoi.png

Duonychus [22]

Gen. et sp. nov

Valid

Kobayashi et al

Late Cretaceous (CenomanianConiacian)

Bayanshiree Formation

Flag of Mongolia.svg  Mongolia

A therizinosaurid theropod. The type species is D. tsogtbaatari.

Duonychus tsogtbaatari.png

Dzharacursor [23]

Gen. et comb. nov

Averianov & Sues

Late Cretaceous (Turonian)

Bissekty Formation

Flag of Uzbekistan.svg  Uzbekistan

An ornithomimid theropod. The type species is "Archaeornithomimus" bissektensis Nesov (1995).

Dzharacursor bissektensis.png

Emiliasaura [24]

Gen. et sp. nov

Valid

Coria et al.

Early Cretaceous (Valanginian)

Mulichinco Formation

Flag of Argentina.svg  Argentina

An ornithopod belonging to the group Rhabdodontomorpha. The type species is E. alessandrii. Announced in 2024; the final article version was published in 2025.

Emiliasaura alessandrii.png
Huadanosaurus [25]

Gen. et sp. nov

Qiu et al.

Early Cretaceous (Barremian)

Yixian Formation

Flag of the People's Republic of China.svg  China

A compsognathid-like theropod belonging to the group Sinosauropterygidae. The type species is H. sinensis.

Huadanosaurus sinensis.png

Mexidracon [26]

Gen. et sp. nov

In press

Serrano-Brañas et al.

Late Cretaceous (Campanian)

Cerro del Pueblo Formation

Flag of Mexico.svg  Mexico

An ornithomimid theropod. The type species is M. longimanus.

Mexidracon longimanus.png

Petrustitan [27]

Gen. et comb. nov

Díez Díaz et al.

Late Cretaceous (Maastrichtian)

Sînpetru Formation

Flag of Romania.svg  Romania

A titanosaur sauropod. The type species is "Magyarosaurus" hungaricus Huene (1932).

Petrustitan hungaricus.png

Qianjiangsaurus [28]

Gen. et sp. nov

Valid

Dai et al.

Late Cretaceous

Zhengyang Formation

Flag of the People's Republic of China.svg  China

An early-diverging hadrosauromorph. The type species is Q. changshengi. Announced in 2024; the final article version was published in 2025.

Qianjiangsaurus changshengi.png

Sinosauropteryx lingyuanensis [25]

Sp. nov

Qiu et al.

Early Cretaceous (Barremian)

Yixian Formation

Flag of the People's Republic of China.svg  China

A compsognathid-like theropod; a species of Sinosauropteryx .

Sinosauropteryx lingyuanensis.png

Tameryraptor [29]

Gen. et sp. nov

Valid

Kellermann, Cuesta & Rauhut

Late Cretaceous (Cenomanian)

Bahariya Formation

Flag of Egypt.svg  Egypt

A carcharodontosaurid theropod. The type species is T. markgrafi.

Tameryraptor markgrafi.png

Uriash [27]

Gen. et sp. nov

Díez Díaz et al.

Late Cretaceous (Maastrichtian)

Densuș-Ciula Formation

Flag of Romania.svg  Romania

A titanosaur sauropod. The type species is U. kadici.

Uriash kadici.png

Xingxiulong yueorum [30]

Sp. nov

Chen et al.

Early Jurassic

Lufeng Formation

Flag of the People's Republic of China.svg  China

A massopodan sauropodomorph; a species of Xingxiulong .

Xingxiulong yueorum.png

Yuanyanglong [31]

Gen. et sp. nov

Valid

Hao et al.

Early Cretaceous

Miaogou Formation

Flag of the People's Republic of China.svg  China

An oviraptorosaurian theropod. The type species is Y. bainian. Announced in 2024; the final article version was published in 2025.

Yuanyanglong bainian.png

General non-avian dinosaur research

Saurischian research

Theropod research

  • A study on the shape and growth of snouts and beaks of extinct theropods and extant birds, providing evidence of a conserved growth pattern of the rostrum throughout the evolutionary history of theropods, is published by Garland et al. (2025). [43]
  • Marques et al. (2025) compare the performance of different machine learning models used for identification of isolated theropod teeth. [44]
  • Piñuela et al. (2025) report the discovery of a theropod footprint preserved with a detached sandstone undertrack from the Upper Jurassic Lastres Formation (Spain), providing evidence of foot movement through the sediment and evidence of changes of footprint morphology at different levels of sediment depth, with some of the successive footprint outlines showing similarities to footprints of members of different dinosaur groups; the authors also reevaluate the type series of the ichnotaxon Iguanodontipus , and argue that some of the studied footprints might have been produced by a theropod. [45]
  • A study on bone histology of Ceratosaurus , providing evidence of faster growth rate than in Late Cretaceous members of Ceratosauria, is published by Sombathy, O'Connor & D'Emic (2025). [46]
  • A study on the maxillary shape of abelisaurids and its relation to feeding ecology is published by Seculi Pereyra et al. (2025), who find evidence of morphological similarities between the maxillae of Spectrovenator and Late Cretaceous abelisaurids, interpreted as likely to be specialist hunters holding and killing prey with their jaws. [47]
  • Redescription of the anatomy of the appendicular skeleton of Piatnitzkysaurus floresi and a study on the phylogenetic affinities of this species is published by Pradelli, Pol & Ezcurra (2025). [48]
  • Isasmendi et al. (2025) describe new fossil material of early-branching tetanurans and baryonychine spinosaurids from the Lower Cretaceous Golmayo Formation (Spain), including a large-bodied baryonychine from the Zorralbo I locality. [49]
  • Evidence indicating that oxygen isotope composition in tooth dentine of Spinosaurus aegyptiacus can be used as a proxy for environmental reconstructions is presented by Liu et al. (2025), who record oxygen isotope variability in the dentine of the studied theropod, interpreted as likely reflecting seasonal environmental changes. [50]
  • Kotevski et al. (2025) describe new fossil material of theropods from the Lower Cretaceous Strzelecki Group and Eumeralla Formation (Australia), including the first carcharodontosaurian fossils from Australia, bones of large-bodied megaraptorids and a tibia of a member of Unenlagiinae. [51]
  • Calvo et al. (2025) report the first discovery of the humerus of an adult specimen of Megaraptor namunhuaiquii from the Upper Cretaceous Portezuelo Formation (Argentina), and interpret its anatomy as indicating that M. namunhuaiquii and Gualicho shinyae were not closely related. [52]
  • A study on the evolution of adaptations to cursoriality in the hindlimbs of theropod dinosaurs and on the origin of arctometatarsus in members of Coelurosauria is published by Kubo & Kobayashi (2025) [53]
  • Scherer (2025) reeavulates evidence for anagenesis in tyrannosaurine tyrannosaurids, and recovers species belonging to the genus Daspletosaurus as forming an evolutionary grade within Tyrannosaurinae, but does not recover Daspletosaurus as a direct ancestor of Tyrannosaurini. [54]
  • Warner-Cowgill et al. (2025) describe a new specimen of Daspletosaurus from the Judith River Formation (Montana, United States), report evidence of the presence of a combination of anatomical features unknown in other members of the genus, and interpret the anatomy of the specimen as weakening the case that D. wilsoni and D. torosus are distinct species. [55]
  • Meso et al. (2025) revise alvarezsaurid fossils from the Salitral Ojo de Agua locality (Allen Formation; Río Negro Province, Argentina) described by Salgado et al. (2009) [56] and an alvarezsaurid femur from the same locality originally described as an ornithopod femur by Coria, Cambiaso & Salgado (2007), [57] describe additional alvarezsaurid material from this locality, and interpret the studied fossils as likely bones of Bonapartenykus ultimus , providing new information on the body plan of members of Patagonykinae. [58]
  • Evidence indicating that digit loss and reduction of the rest of the forelimb in members of Oviraptorosauria were independent changes resulting from different evolutionary processes is presented by Mead, Funston & Brusatte (2025). [59]
  • Zhu et al. (2025) report the discovery of clutch of elongatoolithid eggs from the Upper Cretaceous Qiupa Formation (China), possibly produced by Yulong mini . [60]
  • Foster, Norell & Balanoff (2025) describe two new specimens of Conchoraptor gracilis from the Barun Goyot Formation (Mongolia), present an updated diagnosis for Conchoraptor and differentiate C. gracilis from both Heyuannia yanshini and Khaan mckennai . [61]

Sauropodomorph research

  • Peyre de Fabrègues et al. (2025) describe new fossil material of Leyesaurus marayensis from the Balde de Leyes Formation (Argentina) and revise the anatomy of the holotype specimen of this species, identifying the holotype as a likely juvenile specimen. [62]
  • Toefy, Krupandan & Chinsamy (2025) study the bone histology of two sauropodiform specimens and one early sauropod from the Elliot Formation (South Africa), providing evidence that the three studied specimens underwent rapid growth but differed in the duration of uninterrupted growth, and argue that the change of growth dynamics throughout the evolutionary history of sauropodomorphs was more complex than a simple progression from slow, interrupted growth to fast, uinterrupted growth. [63]
  • Description of the anatomy of the appendicular skeleton of Bagualia alba is published by Gomez et al. (2025), who also study morphological diversity of sauropodomorphs throughout their evolutionary history, and report evidence of shifts in morphospace occupation during the Jurassic related to the diversification of early sauropods and extinction of other sauropodomorphs, as well as to subsequent diversification of Neosauropoda. [64]
  • Saleiro & Tschopp (2025) describe a previously unstudied collection of sauropod teeth from the Upper Jurassic strata in Portugal, identified as belonging to members of Turiasauria, Flagellicaudata, Camarasauridae and Titanosauriformes. [65]
  • A revision of the known material assigned to the genus Haplocanthosaurus is published by Boisvert et al. (2025). [66]
  • A study on the morphology of teeth, their replacement process and possible feeding ecology of Bajadasaurus pronuspinax is published by Garderes (2025). [67]
  • Lerzo & Gallina (2025) redescribe the left ilium of Cathartesaura anaerobica , and interpret its anatomy as consistent with the invasion of the space within the ilium by parts of the abdominal air sac that provided resistance to the thin ilium. [68]
  • Redescription of Liaoningotitan sinensis is published by Shan (2025). [69]
  • Fossil material of lithostrotian titanosaurs assigned to two morphotypes, including caudal vertebrae preserved with rare pathological features, is described from the Upper Cretaceous Cambambe Basin (Brazil) by Lacerda et al. (2025). [70]

Ornithischian research

Thyreophoran research

  • Rivera-Sylva et al. (2025) describe new fossil material of members of Ankylosauria from the Upper Cretaceous strata in Coahuila (Mexico), including fossils from the Maastrichtian Cañon del Tule Formation representing the youngest records of the group from Mexico reported to date. [72]
  • Álvarez Nogueira et al. (2025) report fragmentary remains of a possible parankylosaurian from the Allen Formation (Argentina), likely representing a taxon distinct from the coeval Patagopelta . [73]

Cerapod research

Birds

New bird taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Apus boanoi [84]

Sp. nov

Pavia et al.

Pliocene

Langebaanweg

Flag of South Africa.svg  South Africa

A swift, a species of Apus

Australarus [85]

Gen. et sp. nov

De Pietri et al.

Miocene

Bannockburn Formation

Flag of New Zealand.svg  New Zealand

A member of the family Laridae. The type species is A. bakeri.

Baminornis [86]

Gen. et sp. nov

Valid

Chen et al.

Late Jurassic (Tithonian)

Nanyuan Formation

Flag of the People's Republic of China.svg  China

An early avialan bearing a pygostyle. The type species is B. zhenghensis.

Gracanicanetta [87]

Gen. et sp. nov

Valid

Bocheński et al.

Miocene (Langhian)

Flag of Bosnia and Herzegovina.svg  Bosnia and Herzegovina

A duck. The type species is G. happi.

Hunucornis [88]

Gen. et sp. nov

Agnolín et al.

Miocene

Las Flores Formation

Flag of Argentina.svg  Argentina

A grebe. Genus includes new species H. huayanen.

Miolarus [85]

Gen. et sp. nov

De Pietri et al.

Miocene

Bannockburn Formation

Flag of New Zealand.svg  New Zealand

A member of the family Laridae. The type species is M. rectirostrum.

Novavis [89]

Gen. et sp. nov

In press

O'Connor et al.

Early Cretaceous

Xiagou Formation

Flag of the People's Republic of China.svg  China

A enantiornithean. The type species is N. pubisculata.

Palaelodus haroldocontii [88]

Sp. nov

Agnolín et al.

Miocene

Las Flores Formation

Flag of Argentina.svg  Argentina

? Pseudocrypturus danielsi [90]

Sp. nov

Valid

Mayr & Kitchener

Eocene (Ypresian)

London Clay

Flag of the United Kingdom.svg  United Kingdom

A member of the family Lithornithidae; a species of Pseudocrypturus.

? Pseudocrypturus gracilipes [90]

Sp. nov

Valid

Mayr & Kitchener

Eocene (Ypresian)

London Clay

Flag of the United Kingdom.svg  United Kingdom

A member of the family Lithornithidae; a species of Pseudocrypturus.

Shuilingornis [91]

Gen. et sp. nov

Valid

Wang et al.

Early Cretaceous

Jiufotang Formation

Flag of the People's Republic of China.svg  China

A euornithean in the family Gansuidae. The type species is S. angelai. Announced in 2024; the final article version was published in 2025.

Shuilingornis angelai.png

Zqueheanas [88]

Gen. et sp. nov

Agnolín et al.

Miocene

Las Flores Formation

Flag of Argentina.svg  Argentina

A duck belonging to the subfamily Tadorninae. Genus includes new species Z. hebe.

Avian research

Pterosaurs

New pterosaur taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Darwinopterus camposi [109]

Sp. nov

Valid

Cheng et al.

Jurassic

Tiaojishan Formation

Flag of the People's Republic of China.svg  China

Reconstruction of the skull and lower jaw of Darwinopterus camposi.jpg

Garudapterus [110]

Gen. et sp. nov

Manitkoon et al.

Early Cretaceous

Khorat Group

Flag of Thailand.svg  Thailand

A member of the family Ctenochasmatidae belonging to the subfamily Gnathosaurinae. The type species is G. buffetauti.

Infernodrakon [111]

Gen. et sp. nov

Thomas et al.

Late Cretaceous (Maastrichtian)

Hell Creek Formation

Flag of the United States.svg  United States
(Flag of Montana.svg  Montana)

A member of the family Azhdarchidae. The type species is I. hastacollis.

Infernodrakon.png

Nipponopterus [112]

Gen. et sp. nov

In press

Zhou et al.

Late Cretaceous

Mifune Group

Flag of Japan.svg  Japan

A member of the family Azhdarchidae. The type species is N. mifunensis. Announced in 2024; the final article version was published in 2025.

Nipponopterus Skeletal.svg

Pterosaur research

Other archosaurs

Other new archosaur taxa

NameNoveltyStatusAuthorsAgeType localityCountryNotesImages

Gondwanax [121]

Gen. et sp. nov

Valid

Müller

MiddleLate Triassic (Ladinian–early Carnian)

Pinheiros-Chiniquá Sequence of the Santa Maria Supersequence

Flag of Brazil.svg  Brazil

A sulcimentisaurian member of the possibly paraphyletic family Silesauridae. The type species is G. paraisensis. Announced in 2024; the final article version was published in 2025.

Gondwanax paraisensis.png

Other archosaur research

General research

References

  1. Haldar, A.; Ray, S.; Bandyopadhyay, S. (2025). "A new paratypothoracin aetosaur (Archosauria: Pseudosuchia) from the Upper Triassic Dharmaram Formation of India and its biostratigraphic implications". Journal of Vertebrate Paleontology. e2439533. doi:10.1080/02724634.2024.2439533.
  2. Wu, X.-C.; Witmer, L. M.; Chatterjee, S.; Cunningham, D. (2025). "A new crocodylomorph (Pseudosuchia, Crocodylomorpha) from the Upper Triassic of Texas and its phylogenetic relationships". Journal of Vertebrate Paleontology. e2446604. doi:10.1080/02724634.2024.2446604.
  3. Bravo, Gonzalo Gabriel; Pol, Diego; Leardi, Juan Martín; Krause, Javier Marcelo; Nicholl, Cecily S. C.; Rougier, Guillermo; Mannion, Philip D. (2025-03-26). "A new notosuchian crocodyliform from the Early Palaeocene of Patagonia and the survival of a large-bodied terrestrial lineage across the K–Pg mass extinction". Proceedings of the Royal Society B: Biological Sciences. 292 (2043): 20241980. doi:10.1098/rspb.2024.1980.
  4. Carvalho, J. C.; Santos, D. M.; Pinto, R. L.; Santucci, R. M. (2025). "Anatomical description and systematics of a new notosuchian (Mesoeucrocodylia; Crocodyliformes) from the Quiricó Formation, Lower Cretaceous, Sanfranciscana Basin, Brazil". Journal of Vertebrate Paleontology. e2452947. doi:10.1080/02724634.2025.2452947.
  5. Fitch, A. J.; Kammerer, C. F.; Nesbitt, S. J. (2025). "First occurrences of Poposauroidea (Archosauria: Paracrocodylomorpha) from North Carolina expand their geographic range in the Late Triassic". Palaeodiversity. 18 (1): 1–9. doi: 10.18476/pale.v18.a1 .
  6. McDavid, Skye Noin (2025-03-15). "Huenesuchus is an objective synonym of Prestosuchus while 'class-group names' do not exist in and are not regulated by the ICZN: a response to Kischlat". Revista Brasileira de Paleontologia. 27 (4): e20240425. doi:10.4072/rbp.2024.4.0425.
  7. Błaszczeć, P.; Antczak, M. (2025). "The histology and function of the dermal armour of the aetosaur Stagonolepis olenkae Sulej, 2010 (Archosauria, Pseudosuchia) from Krasiejów (SW Poland)". Acta Geologica Polonica. 75 (1). e38. doi: 10.24425/agp.2024.152660 .
  8. Ponce, D. A.; Cerda, I. A.; Desojo, J. B. (2025). "A fast start: Evidence of rapid growth in Trialestes romeri, an early Crocodylomorpha from the Upper Triassic continental beds of Argentina based on osteohistological analyses". Journal of Anatomy. doi:10.1111/joa.14230. PMID   39887998.
  9. Forêt, T.; Aubier, P.; Jouve, S.; Cubo, J. (2025). "Analysing Thalattosuchia palaeobiodiversity through the prism of phylogenetic comparative methods". Palaeontology. 68 (1). e70000. doi: 10.1111/pala.70000 .
  10. Johnson, M. M.; Sachs, S.; Young, M. T.; Abel, P. (2025). "A re-description of the teleosauroid Macrospondylus bollensis (Jaeger, 1828) from the Posidonienschiefer Formation of Germany". PalZ. doi: 10.1007/s12542-024-00712-x .
  11. Sena, M. V. A.; Montefeltro, F. C.; Marinho, T. S.; Langer, M. C.; Fachini, T. S.; Pinheiro, A. E. P.; Machado, A. S.; Lopes, R. T.; Pellarin, R.; Sayao, J. M.; Oliveira, G. R.; Cubo, J. (2025). "Revisiting the aerobic capacity of Notosuchia (Crocodyliformes, Mesoeucrocodylia)". Lethaia. 57 (4): 1–8. doi: 10.18261/let.57.4.6 .
  12. Navarro, T. G.; Cerda, I. A.; Filippi, L. S.; Pol, D. (2025). "Life history and growth dynamics of a peirosaurid crocodylomorph (Mesoeucrocodylia; Notosuchia) from the Late Cretaceous of Argentina inferred from its bone histology". Journal of Anatomy. doi:10.1111/joa.14182. PMID   39846495.
  13. Kuzmin, I. T.; Sichinava, E. A.; Mazur, E. V.; Gombolevskiy, V. A.; Sennikov, A. G.; Skutschas, P. P. (2025). "Neurocranial anatomy of Paralligator (Neosuchia: Paralligatoridae) from the Upper Cretaceous of Mongolia". Zoological Journal of the Linnean Society. 203 (1). zlae177. doi:10.1093/zoolinnean/zlae177.
  14. Lessner, E. J.; Petermann, H.; Lyson, T. R. (2025). "First record of Borealosuchus sternbergii from the lower Paleocene Denver Formation (lower Danian), Colorado (Denver Basin)". Journal of Vertebrate Paleontology. e2434214. doi:10.1080/02724634.2024.2434214.
  15. Hoffman, D. K.; Goldsmith, E. R.; Houssaye, A.; Maidment, S. C. R.; Felice, R. N.; Mannion, P. D. (2025). "Evolution of growth strategy in alligators and caimans informed by osteohistology of the late Eocene early-diverging alligatoroid crocodylian Diplocynodon hantoniensis". Journal of Anatomy. doi: 10.1111/joa.14231 . PMID   39924872.
  16. Serrano-Martínez, A.; Luján, À. H.; García-Pérez, Á.; Fortuny, J. (2025). "New data on the inner skull cavities of Diplocynodon tormis (Crocodylia, Diplocynodontinae) from the Duero Basin (Iberian Peninsula, Spain)". Fossil Record. 28 (1): 67–77. doi: 10.3897/fr.28.133743 .
  17. Pligersdorffer, C. C.; Burke, P. M. J.; Mannion, P. D. (2025). "Evaluation of the endocranial anatomy of the early Paleogene north African gavialoid crocodylian Argochampsa krebsi and evolutionary implications for adaptation to salinity tolerance in marine crocodyliforms". Journal of Anatomy. doi: 10.1111/joa.14213 . PMID   39814549.
  18. El-Degwi, E. S.; AbdelGawad, M.; Radwaan, S. E.; Sliem, R. E.; Sileem, A.; Abd Elhady, S. I. (2025). "Evolutionary trend of the broad-snouted crocodile from the Eocene, Early Miocene and recent ones from Egypt". Scientific Reports. 15. 9159. doi: 10.1038/s41598-025-91167-w .
  19. Lovelace, David M; Kufner, Aaron M; Fitch, Adam J; Curry Rogers, Kristina; Schmitz, Mark; Schwartz, Darin M; LeClair-Diaz, Amanda; St.Clair, Lynette; Mann, Joshua; Teran, Reba (2025-01-01). "Rethinking dinosaur origins: oldest known equatorial dinosaur-bearing assemblage (mid-late Carnian Popo Agie FM, Wyoming, USA)". Zoological Journal of the Linnean Society. 203 (1): zlae153. doi:10.1093/zoolinnean/zlae153. ISSN   0024-4082.
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