Shuilingornis Temporal range: Early Cretaceous, | |
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Reconstructed skeleton | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Family: | † Gansuidae |
Genus: | † Shuilingornis Wang et al., 2024 |
Species: | †S. angelai |
Binomial name | |
†Shuilingornis angelai Wang et al., 2024 | |
Shuilingornis (meaning "pretty and vivid bird") is an extinct genus of gansuid euornithean birds from the Early Cretaceous Jiufotang Formation of China. The genus contains a single species, S. angelai, known from a nearly complete articulated skeleton. As a member of the Gansuidae, Shuilingornis represents one of the earliest known birds known to demonstrate semi-aquatic adaptations. [1]
The Shuilingornis holotype specimen, LY2022JZ3002, was discovered in sediments of the Jiufotang Formation ('Lamadong locality') in Jianchang County of Huludao City, Liaoning Province, China. The specimen is a nearly complete and articulated skeleton, preserved on a single slab. Some feather imprints are visible around the hand of the holotype. An egg-shaped pigmented spot is preserved in the abdominal region, indicating some form of soft tissue. Similarly, a dark spot observed in the orbital region likely represents traces of the eye. [1]
In 2024, Wang et al. described Shuilingornis angelai as a new genus and species of gansuid avialans based on these fossil remains. The generic name, Shuilingornis, combines the Chinese word "shuiling", meaning "pretty and vivid", with the Greek word "ornis", meaning "bird". The specific name, angelai, honors Piero Angela, an Italian science communicator and journalist. [1]
Based on the length of the tibiotarsus, Shuilingornis was smaller than other members of the Gansuidae, 70% the size of Gansus , and 88% the size of Changzuiornis . Bone histology of the preserved ulna, indicates that the Shuilingornis holotype was a young adult when it died. [1]
In their phylogenetic analyses, Wang et al. (2024) recovered Shuilingornis as a member of the euornithean clade Gansuidae, closely related to Changzuiornis. Other members of this clade include Gansus, Iteravis , Khinganornis , and possibly Hollanda . Their results are displayed in the cladogram below: [1]
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Gansus is a genus of aquatic birds that lived during the Aptian age of the Early Cretaceous (Aptian-Albian) period in what are now Gansu and Liaoning provinces, western China. The rock layers from which their fossils have been recovered are dated to 120 million years ago. It was first described in 1984 on the basis of an isolated left leg. It is the oldest-known member of the Ornithurae, the group which includes modern birds (Neornithes) and extinct related groups, such as Ichthyornis and Hesperornithes.
Gansuidae is an extinct family of euornithean birds known from the Early Cretaceous of China and possibly the Late Cretaceous of Mongolia.
Teviornis is an extinct genus of presbyornithid which lived during the Maastrichtian stage, around 70 million years ago. One species has been described, T. gobiensis. It is the oldest known neognath and its fossils are collected from the Nemegt Formation of Mongolia.
Chaoyangia is an extinct genus of euornithean birds, containing the single species Chaoyangia beishanensis. This species is known from a single fossil specimen consisting of a partial skeleton including vertebra, ribs, hips, and upper legs. The specimen was discovered in the Jiufotang Formation near the city of Chaoyang in Liaoning province, China. This rock formation has been dated to the Aptian age of the Early Cretaceous period, 120 million years ago.
Zhongjianornis is a genus of beaked, pigeon-sized birds from the early Cretaceous period of China. It is known from one fossil found at Jianchang, Liaoning Province, in rocks of the Jiufotang Formation, representing the type species Zhongjianornis yangi.
Rapaxavis a genus of enantiornithine bird. It has been found in the Jiufotang Formation in Liaoning, People's Republic of China.
Parabohaiornis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Parabohaiornis martini.
Longusunguis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Longusunguis kurochkini.
Chuanqilong is a monospecific genus of basal ankylosaurid dinosaur from the Liaoning Province, China that lived during the Early Cretaceous in what is now the Jiufotang Formation. The type and only species, Chuanqilong chaoyangensis, is known from a nearly complete skeleton with a skull of a juvenile individual. It was described in 2014 by Fenglu Han, Wenjie Zheng, Dongyu Hu, Xing Xu, and Paul M. Barrett. Chuanqilong shows many similarities with Liaoningosaurus and may represent a later ontogenetic stage of the taxon.
Zhenyuanlong is a genus of dromaeosaurid dinosaur from the Yixian Formation of Liaoning, China. It lived during the Aptian age of the early Cretaceous period, approximately 125 million years ago. It is known from a single specimen belonging to the species Zhenyuanlong suni. This type specimen preserved a nearly complete skeleton that contains traces of feathers, including long tail feathers and large wings. In addition to further complicating diversity of Liaoning dromaeosaurids, this specimen provides the first direct evidence of well-developed pennaceous feathers in a large, non-flying dromaeosaur, raising the question of what function such wings would serve.
Forfexopterus is a genus of ctenochasmatid pterosaur from the Early Cretaceous Jiufotang Formation in China. It contains a single species, F. jeholensis, named from a mostly complete skeleton by Shunxing Jiang and colleagues in 2016. A second specimen, consisting of a wing, was described in 2020. While the first specimen is larger, it shows signs of being less mature than the second specimen, indicating that the developmental trajectories of Forfexopterus were variable. Like other ctenochasmatids, Forfexopterus had a long, low skull filled with many slender teeth; unlike other members of the group, however, it did not have a spatula-shaped snout tip or crests, and its teeth were more curved. A single characteristic distinguishes Forfexopterus from all other members of the wider group Archaeopterodactyloidea: of the four phalanx bones in its wing finger, the first was shorter than the second but longer than the third.
Changzuiornis is an extinct genus of ornithuromorph bird from the Early Cretaceous of present-day China. It contains a single species, C. ahgmi.
Juehuaornis is an extinct genus of ornithuromorph dinosaurs from the Early Cretaceous of present-day China. It contains a single species, J. zhangi.
Chiappeavis is a genus of enantiornithean bird from Early Cretaceous of northeastern China. The only species is Chiappeavis magnapremaxillo. Chiappeavis is classified within the family Pengornithidae. It is known from a single, almost complete skeleton including feather impressions discovered in the Jiufotang Formation of the Jehol Group. Long feathers formed a fan-shaped tail that was probably employed in flight.
Musivavis is a genus of euenantiornithine bird from the Early Cretaceous (Aptian) Jiufotang Formation of Liaoning Province, China. The genus contains a single species, Musivavis amabilis, known from a nearly complete, articulated skeleton.
Daurlong is an extinct genus of dromaeosaurid dinosaur from the Early Cretaceous (Aptian) Longjiang Formation of China. The genus contains a single species, D. wangi, known from a nearly complete skeleton. Daurlong represents the first described occurrence of a preserved intestinal region in a theropod closely related to birds.
Cratonavis is an extinct genus of pygostylian avialian from the Early Cretaceous Jiufotang Formation of Liaoning Province, China. The genus contains a single species, C. zhui, known from a complete skeleton.
Meilifeilong is an extinct genus of chaoyangopterid pterosaurs from the Early Cretaceous Jiufotang Formation of China. The genus contains two species: M. youhao, known from an almost complete skeleton and a partial skull, and M. sanyainus, originally described as a species of Shenzhoupterus. The Meilifeilong youhao holotype represents the best-preserved and most complete chaoyangopterid discovered so far.
Imparavis is an extinct genus of enantiornithean birds from the Early Cretaceous (Aptian-aged) Jiufotang Formation of Liaoning Province, China. The genus contains a single species, I. attenboroughi, known from a well-preserved skeleton.
Neobohaiornis is an extinct genus of bohaiornithid enantiornithean birds from the Early Cretaceous (Aptian-aged) Jiufotang Formation of Liaoning Province, China. The genus contains a single species, N. lamadongensis, known from a well-preserved skeleton with feather impressions.