Ornithothoracines | |
---|---|
Fossil of an enantiornithean ( Junornis houi ) | |
Southern ground hornbill (Bucorvus leadbeateri) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Clade: | Pygostylia |
Clade: | Ornithothoraces Chiappe & Calvo, 1994 |
Subgroups | |
Ornithothoraces is a group of avialan dinosaurs that includes all enantiornithes ("opposite birds") and the euornithes ("true birds"), which includes modern birds and their closest ancestors. The name Ornithothoraces means "bird thoraxes". This refers to the modern, highly advanced anatomy of the thorax that gave the ornithothoracines superior flight capability compared with more primitive avialans. This anatomy includes a large, keeled breastbone, elongated coracoids and a modified glenoid joint in the shoulder, and a semi-rigid rib cage. In spite of this at least the sternum seems to have developed convergently rather than being a true homology. [1]
The earliest known members of the group are the enantiornitheans Protopteryx fengningensis , Eopengornis martini , and Cruralispennia multidonta , as well as the euornithine Archaeornithura meemannae , all from the Sichakou Member of the Huajiying Formation in China, which has been dated to 130.7 million years old. [2] At least one other enantiornithean, Noguerornis gonzalezi , may be even older, at up to 145.5 million years ago, though its exact age is uncertain. [3]
In 1994, Chiappe and Calvo established a phylogenetic definition of the group. They defend Ornithothoraces as a node-based clade, the common ancestor of Iberomesornis romerali and modern birds, and all of its other descendants. [4] In 1998, Paul Sereno defined Ornithothoraces in the same way, but used Sinornis santensis instead of Iberomesornis romerali. [5]
The cladogram below follows the results of a phylogenetic analysis by Wang et al., 2016: [6]
Ornithothoraces | |
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Yanornis is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.
Jeholornis is a genus of avialan dinosaurs that lived between approximately 122 and 120 million years ago during the early Cretaceous Period in China. Fossil Jeholornis were first discovered in the Jiufotang Formation in Hebei Province, China and additional specimens have been found in the older Yixian Formation.
Nanantius is a genus of extinct enantiornithean avialan known from the Early Cretaceous of Australia.
Longipteryx is a genus of prehistoric bird which lived during the Early Cretaceous. It contains a single species, Longipteryx chaoyangensis. Its remains have been recovered from the Jiufotang Formation at Chaoyang in Liaoning Province, China. Apart from the holotype IVPP V 12325 - a fine and nearly complete skeleton — another entire skeleton and some isolated bones are known to date.
Eoalulavis is a monotypic genus of enantiornithean bird that lived during the Barremian, in the Lower Cretaceous around 125 million years ago. The only known species is Eoalulavis hoyasi.
Cathayornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known definitively from only one species, Cathayornis yandica, one of the first Enantiornithes found in China. Several additional species were once incorrectly classified as Cathayornis, and have since been reclassified or regarded as nomina dubia.
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces.
The Jiufotang Formation is an Early Cretaceous geological formation in Chaoyang, Liaoning which has yielded fossils of feathered dinosaurs, primitive birds, pterosaurs, and other organisms. It is a member of the Jehol group. The exact age of the Jiufotang has been debated for years, with estimates ranging from the Late Jurassic to the Early Cretaceous. New uranium-lead dates reveal the formation is deposited in the Aptian stage of the Early Cretaceous. Fossils of Microraptor and Jeholornis are from the Jiufotang.
Protopteryx is an extinct bird and possibly the basalmost enantiornithean, from the Cretaceous period. The type species is P. fengningensis. It was first discovered in the Sichakou Member of the Yixian Formation or Huajiying Formation of Hebei Province, northern China, dating from 131 Ma ago. Protopteryx has been found in the Daibeigou formation, as well. The name Protopteryx means "primitive feather": "proto-" meaning "the first of" and "-pteryx" meaning "feather" or "wing." The name comes from the fact that Protopteryx feathers are more primitive than those of modern birds, such as the two elongated tail feathers that lack barbs and rami.
Avisauridae is a family of extinct enantiornithine dinosaurs from the Cretaceous period, distinguished by several features of their ankle bones. Depending on the definition used, Avisauridae is either a broad and widespread group of advanced enantiornithines, or a small family within that group, restricted to species from the Late Cretaceous of North and South America.
Shanweiniao is a genus of long-snouted enantiornithean birds from Early Cretaceous China. One species is known, Shanweiniao cooperorum. There is one known fossil, a slab and counterslab. The fossil is in the collection of the Dalian Natural History Museum, and has accession number DNHM D1878/1 and DNHM1878/2. It was collected from the Lower Cretaceous Dawangzhengzi Beds, middle Yixian Formation, from Lingyuan in the Liaoning Province, China.
Hongshanornithidae is an extinct group of early ornithuromorph birds from the early Cretaceous period of China. It includes the genera Hongshanornis and Tianyuornis from the Yixian Formation of Inner Mongolia, Longicrusavis from the Yixian Formation of Liaoning Province, Parahongshanornis from the Jiufotang Formation of Liaoning Province, and Archaeornithura, the oldest known member, from the Huajiying Formation of Hebei Province.
Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.
Euornithes is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to Sinornis.
Pengornithidae is a group of early enantiornithines from the early Cretaceous Period of China, with the putative member Falcatakely possibly extending this clade's range into the Late Cretaceous of Madagascar, and several putative pengornithids also hail from this formation. Specimens of these animals have been found both in the Huajiying Formation and Jiufotang Formation of Liaoning and Hebei provinces, dating from the Hauterivian age to the Aptian age.
Cruralispennia is an extinct genus of enantiornithean bird. The only known specimen of Cruralispennia was discovered in the Early Cretaceous Huajiying Formation of China and formally described in 2017. The type species of Cruralispennia is Cruralispennia multidonta. The generic name is Latin for "shin feather", while the specific name means "many-toothed". The holotype of Cruralispennia is IVPP 21711, a semi-articulated partial skeleton surrounded by the remains of carbonized feathers.
Chongmingia is a genus of basal avialan belonging to Pygostylia that lived during the Aptian. It was found in the Jiufotang Formation in Chaoyang, China, and was described by Wang et al., 2016. The name comes from the word Chongming, referring to a Chinese mythological bird, and the specific epithet is in honor of Mr. Xiaoting Zheng.