Yixianosaurus Temporal range: Early Cretaceous, | |
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A diagram of preserved elements of IVPP V12638 | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Anchiornithidae |
Genus: | † Yixianosaurus Xu & Wang, 2003 |
Species: | †Y. longimanus |
Binomial name | |
†Yixianosaurus longimanus Xu & Wang, 2003 | |
Yixianosaurus (meaning "Yixian lizard") is a maniraptoran theropod dinosaur genus from the Early Cretaceous of China.
The type species, Yixianosaurus longimanus, was formally named and described by Xu Xing and Wang Xiaolin in 2003. Its partial skeleton was discovered in 2001, in Liaoning at Wangjiagou in northeastern China. The generic name refers to the Yixian Formation. The specific name means "with a long hand" from Latin longus, "long", and manus, "hand". [1]
Yixianosaurus is known only from a single specimen, holotype IVPP V12638, which likely derived from the Dawangzhangzi Bed (early Aptian stage, 122 million years ago). [2] It is a compression fossil, viewed from behind and preserved on a single slab that has been sawed into several pieces. It consists of the shoulder girdle and a pair of fossilized arms complete with fossilized feathers, some ribs, and gastralia. Yixianosaurus has a very long hand, 140% of the length of the 89 millimetres (3.5 inches) humerus. The second finger is the longest. The fingers bear large and recurved claws. The feathers are not preserved well enough to show a specific structure, but they appear similar to the contour feathers of some Yixian Formation birds. [1] The large hands could have served in catching prey or assisted in climbing. The total body length has been estimated at 1 meter (3.3 feet), the weight at 1 kilogram (2.2 pounds). [3] Xu et al. (2013) suggested that the presence of large pennaceous feathers on parts of the forelimb strongly supports that Yixianosaurus was adapted for limited aerial locomotion. [4]
The describers considered the exact placement of Yixianosaurus within Maniraptora to be uncertain, but because the hand length resembled that of another feathered dinosaur, Epidendrosaurus (now Scansoriopteryx ), they suggested it was a close relative of the Scansoriopterygidae. Other researchers have suggested the specimen may have come from a dromaeosaurid. Subsequent analyses were divided on whether is it is more primitive and outside the clade Eumaniraptora – this would mean that advanced characteristics such as the long hands and short arms evolved independently in this species [5] – or a basal member of the more advanced Paraves. [4] In a 2017 re-evaluation of the Harlem Archaeopteryx specimen as an anchiornithid called Ostromia , Yixianosaurus is found to be the most basal paravian. [6] However, two other studies published the same year argued that Yixianosaurus was most closely related to Xiaotingia , with both genera being either relatives of scansoriopterygids [7] or anchiornithids. [8]
Caudipteryx is a genus of small oviraptorosaur dinosaurs that lived in Asia during the Early Cretaceous, around 124.6 million years ago. They were feathered and extremely birdlike in their overall appearance, to the point that some paleontologists suggested it was a bird. Two species have been described: C. zoui, in 1998, and C. dongi, in 2000.
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, Oviraptorosauria, and Therizinosauria. Ornitholestes and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period, and survive today as living birds.
Coelurosauria is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Beipiaosaurus is a genus of therizinosauroid theropod dinosaurs that lived in Asia during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the discovery of Yutyrannus, Beipiaosaurus were among the heaviest dinosaurs known from direct evidence to be feathered. Beipiaosaurus is known from three reported specimens. Numerous impressions of feather structures were preserved that allowed researchers to determine the feathering color which turned out to be brownish.
Sinornithosaurus is a genus of feathered dromaeosaurid dinosaur from the early Cretaceous Period of the Yixian Formation in what is now China. It was the fifth non–avian feathered dinosaur genus discovered by 1999. The original specimen was collected from the Sihetun locality of western Liaoning. It was found in the Jianshangou beds of the Yixian Formation, dated to 124.5 million years ago. Additional specimens have been found in the younger Dawangzhangzi bed, dating to around 122 million years ago.
Jinfengopteryx is a genus of maniraptoran dinosaur. It was found in the Qiaotou Member of the Huajiying Formation of Hebei Province, China, and is therefore of uncertain age. The Qiaotou Member may correlate with the more well-known Early Cretaceous Yixian Formation, and so probably dates to around 122 Ma ago.
Scansoriopterygidae is an extinct family of climbing and gliding maniraptoran dinosaurs. Scansoriopterygids are known from five well-preserved fossils, representing four species, unearthed in the Tiaojishan Formation fossil beds of Liaoning and Hebei, China.
Jeholornis is a genus of avialan dinosaurs that lived between approximately 122 and 120 million years ago during the early Cretaceous Period in China. Fossil Jeholornis were first discovered in the Jiufotang Formation in Hebei Province, China and additional specimens have been found in the older Yixian Formation.
Graciliraptor is a genus of theropod dinosaur from the early Cretaceous Period. It is a microraptorine dromaeosaurid.
Caudipteridae is an extinct family of oviraptorosaurian dinosaurs known from the Early Cretaceous of China. Found in the Yixian and Jiufotang Formations, the group existed between 125 and 120 million years ago. Distinguishing characteristics of this group have been indicated as including a unique dagger-shaped pygostyle. In 2015, the group was defined as "the most inclusive clade containing Caudipteryx zoui but not Oviraptor philoceratops and Caenagnathus collinsi".
Dongbeititan is a genus of sauropod dinosaur from the Early Cretaceous-age Yixian Formation of Beipiao, Liaoning, China. It is based on holotype DNHM D2867, a partial postcranial skeleton including bones from the limbs, shoulder and pelvic girdles, and vertebrae, which was described in 2007. Its describers suggested it was as a basal titanosauriform, not as derived as Gobititan or Jiutaisaurus, but more derived than Euhelopus, Fusuisaurus, and Huanghetitan. The type species is D. dongi, and it is the first named sauropod from the Yixian Formation, which is part of the well-known Jehol Group. The genus name refers to the region Dongbei and to Greek titan, "giant". The specific name honours the Chinese paleontologist Dong Zhiming. Like other sauropods, Dongbeititan would have been a large quadrupedal herbivore.
Zhongornis is a genus of primitive maniraptoran dinosaurs that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008.
The Yixian Formation is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans about 1.6 million years during the early Cretaceous period. It is known for its fossils, listed below.
Xiaotingia is a genus of anchiornithid theropod dinosaur from Middle Jurassic or early Late Jurassic deposits of western Liaoning, China, containing a single species, Xiaotingia zhengi.
Aurornis is an extinct genus of anchiornithid theropod dinosaurs from the Jurassic period of China. The genus Aurornis contains a single known species, Aurornis xui. Aurornis xui may be the most basal ("primitive") avialan dinosaur known to date, and it is one of the earliest avialans found to date. The fossil evidence for the animal pre-dates that of Archaeopteryx lithographica, often considered the earliest bird species, by about 10 million years.
Zhongjianosaurus is a genus of dromaeosaurid belonging to the Microraptoria. Believed to hail from the Yixian Formation, specifically the middle of the Jehol Biota, it is the smallest known microraptorine thus far discovered and one of the smallest non-avian theropod dinosaurs.
Jianianhualong is a genus of troodontid theropod dinosaur from the Early Cretaceous of China. It contains a single species, Jianianhualong tengi, named in 2017 by Xu Xing and colleagues based on an articulated skeleton preserving feathers. The feathers at the middle of the tail of Jianianhualong are asymmetric, being the first record of asymmetrical feathers among the troodontids. Despite aerodynamic differences from the flight feathers of modern birds, the feathers in the tail vane of Jianianhualong could have functioned in drag reduction whilst the animal was moving. The discovery of Jianianhualong supports the notion that asymmetrical feathers appeared early in the evolutionary history of the Paraves.
Anchiornithidae is a family of small paravian dinosaurs. Anchiornithids have been classified at varying positions in the paravian tree, with some scientists classifying them as a distinct family, a basal subfamily of Troodontidae, members of Archaeopterygidae, or an assemblage of dinosaurs that are an evolutionary grade within Avialae or Paraves.