Jianchangosaurus Temporal range: Early Cretaceous, ~ | |
---|---|
![]() | |
Holotype specimen | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | † Therizinosauria |
Genus: | † Jianchangosaurus Pu et al. 2013 |
Type species | |
Jianchangosaurus yixianensis Pu et al. 2013 |
Jianchangosaurus ("lizard from Jianchang County") is a genus of therizinosaurian dinosaur that lived approximately 126 million years ago during the early part of the Cretaceous Period from the Yixian Formation in what is now China. The type specimen, a juvenile, was discovered in Jianchang County, in the western part of Liaoning Province. It was described in 2013 by a team of palaeontologists from the Henan Museum, who determined that it was among the most basal therizinosaurs.
Based on the type specimen, Jianchangosaurus was a small, lightly built, bipedal, ground-dwelling herbivore, which could grow up to an estimated 2 m (6.6 ft) long and was 1 m (3.3 ft) high at the hips. It probably had a body mass of around 26 kg (57 lb). In spite of its basal position, Jianchangosaurus already bore many of the hallmarks of therizinosaur anatomy, such as a beaked premaxilla, and the shape of the pubic boot. Jianchangosaurus' tibia was about half as long again as the femur, suggesting cursorial (running) habits.
The holotype of Jianchangosaurus (41HIII-0308A) is the nearly complete skeleton of a single juvenile, missing only the distal (far) portion of the tail, and some minor elements. It was discovered in Jianchang County, in the western part of Liaoning Province. The strata it was recovered from belong to the Yixian Formation, a Lagerstätte known for its vertebrate fossils. It was purchased by the Henan Geological Museum (now the Henan Museum), who prepared it, unintentionally repositioning some of the elements in the process. In 2013, Hanyong Pu, Yoshitsugu Kobayashi, Junchang Lu, Li Xu, Yanhua Wu, Huali Chang, Jiming Zhang, and Songhai Jia published a paper describing 41HIII-0308A. They designating as it the holotype of a new genus and species of basal therizinosaur, Jianchangosaurus yixianensis. The genus name is derived from the county where the specimen was found, and the Greek word "sauros" (σαυρος), meaning "lizard". The specific name yixianensis, refers to the Yixian Formation where the specimen was found, and the Latin suffix "-ensis" meaning "originating in". [1]
Jianchangosaurus was estimated to have been 1 m (3.3 ft) tall at the hips and approximately 2 m (6.6 ft) long. [1] Gregory S. Paul estimated the weight of the holotype specimen around 20 kg (44 lb), though expressed uncertainty about its total body length, due to the fact that tail tip is unpreserved. [2] In 2024, he suggested a higher body mass of 26 kg (57 lb) [3]
The skull of Jianchangosaurus' holotype is fairly well-preserved, lacking only a few elements. It measures 23 cm (9.1 in) in height, and 11 cm (4.3 in) in height. The premaxilla and the anterior (front) portion of the maxilla are edentulous, meaning they lacked teeth. The premaxilla is lined with small foramina, suggesting that a beak was present. The dorsal (top) border of the antorbital fenestra is formed by the lacrimal, maxilla, and nasal, with the majority being formed by the latter. The posterior (rear) portion of the nasal is wide transversely (from side to side), as in Falcarius. The jugal does not participate in the antorbital fenestra's margin, unlike many other coelurosaurs. The prefrontals are triangular, with a V-shaped anterodorsal region (at the top and towards the front) which contacts the lacrimal. The condyloid (articular) process of the quadrate, to which the lower jaw was articulated, was rounded. Jianchangosaurus' dentary was triangular in lateral view, with a downturned symphyseal region, as in other therizinosaurs (with the exception of Falcarius). At very front of the dentary was a toothless region called a diastema. [1]
Jianchangosaurus had twenty-seven maxillary teeth, and an unknown number of dentary teeth (probably between twenty-five and twenty-eight). This tooth count is similar to that observed in other therizinosaurs, like Erlikosaurus , Falcarius, and Segnosaurus . The teeth were small and lanceolate (shaped like the tip of a lance). The teeth consistently have three denticles of equal size. In the anterior portion of the maxilla, before the edentulous region, the teeth were more densely packed. The anterior dentary teeth were similarly sized to those further back, unlike in Falcarius. The outside surface of each maxillary tooth was convex, while the outside surface of each dentary tooth was concave. A similar dental morphology is seen in ornithopods and ceratopsians, and likely maximised biting stress during occlusion, allowing Jianchangosaurus to more effectively slice plant fibres. [1]
Ten cervical (neck) vertebrae are preserved in Jianchangosaurus, consisting of the atlas (first cervical vertebra), axis (second cervical vertebra), and all of the following vertebrae. All of the preserved cervical vertebrae are amphicoelous, meaning that they were concave at both ends, and were also highly pneumatised. The postzygapophysis of the atlas was short, with a rounded posterior end. The axis had longer postzygapophyses, which extended posterior to its short neural spine, and were longer than the prezygapophyses. The third to sixth cervicals were longer than those at the front. The neural arches of the middle and posterior cervicals were X-shaped in dorsal view, like other therizinosaurs, and like oviraptorosaurs. The dorsal (back) vertebrae of Jianchangosaurus had longer neural spines than any of the cervicals. The anterior dorsal vertebrae lacked prominent hypapophyses; finer details are difficult to discern, as in the holotype they are obscured by the ribs. The neural spines had rounded tips in the case of the anterior vertebrae, though in the middle to posterior dorsal vertebrae, they transition to a more square shape. At least sixteen gastralia, bones which would have supported the abdominal organs and served as muscle attachment sites, were present. They were spint-like in shape. Like other basal therizinosaurs, Jianchangosaurus had five sacral vertebrae. Eleven caudal (tail) vertebrae are preserved, consisting of the entire caudal column up to the eleventh. The neural spines of the caudal vertebrae were tilted posteriorly, with distal tips split into anterior and posterior alae. [1]
The scapula and coracoid of Jianchangosaurus were unfused, like in other basal therizinosaurs, though this may be due to its nature as a juvenile. The glenoid of each scapula faced posteriorly. The scapular blade was straight, and remains nearly consistent in width for the entirety of its length. This is unlike the condition of other therizinosaurs, which either increased or decreased the width of the scapular blade. The coracoid was similar to that of ornithomimosaurs, having a cemicircular outline with a prominent posterior process. The humerus measured 158.5 mm (6.2 in) in length, slightly shorter than the scapula. Like in other basal therizinosaurs, both ends were moderately expanded. The internal tuberosity was large, and was separated from the humeral head by a depression, similar to in Falcarius. Unlike other therizinosaurs, the entepicondyle was reduced. The ulna was around seventy-seven percent as long as the humerus, a ratio also seen in Falcarius. Its proximal (near) tip was triangular, with a poorly developed olecranon process compared to other therizinosaurs. The radius was straight and lacked a biceps tubercle, again similar to Falcarius. Four distal carpals are preserved, including the semilunate carpal that characterises maniraptorans. All metacarpals are preserved, except for the third right metacarpal. The longest preserved phalanx (digit bone; finger bone, in this case) is the first, slightly shorter than the second metacarpal. The manual unguals (hand claws) were curved, and were narrow transversely. The flexor tubercles of all unguals, to which the flexor tendons attached, were rounded. A groove on the medial surface of each ungual reaches the dorsal surface at its tip. [1]
The pelvis of Jianchangosaurus bore several features that were primitive to Coelurosauia, though absent in more derived therizinosaurs. The ilium was low, with a nearly horizontal dorsal edge, a shallow process before the acetabulum (the preacetabular process), and a deep process after it (postacetabular process). The pubis is shorter than the ischium. It projects anteroventrally. As such, unlike more derived therizinosaurs, it is not opisthopubic, meaning that it was not oriented backwards. The femur was straight, with a cylindrical lesser trochanter that was separated from the greater trochanter by a cleft. The neck of the femoral head is weakly constructed. The fourth trochanter is not known. The tibia is around half as long again as the femur, the highest ratio seen in therizinosaurs; [1] such an adaptation strongly correlates to cursorial habits in dinosaurs. [1] [4] The fibula is narrower than the tibia. It is not clear how many of Jianchangosaurus' metatarsals are real, as most appear to have been restored. Most pedal (foot) phalanges cannot be identified, as most are crushed, and elements from the manus (hand) may be mixed in. Four pedal unguals are known, all with poorly developed flexor tubercules. All of them are recurved and transversely compressed, similar to in Erlikosaurus. [1]
The impressions of a series of wide and unbranched feathers surround the holotype of Jianchangosaurus. Only their distal ends are visible. The feathers, considered elongated broad filamentous feathers (EBFFs), are similar to those of Beipiaosaurus , from the same formation. The describers of Jianchangosaurus speculated that they may have been used for visual display. [1]
Like the primitive Falcarius and Beipiaosaurus, Jianchangosaurus was classified as a basal therizinosaur. Phylogenetic analysis suggests that it is more derived than Falcarius but more primitive than Beipiaosaurus. Jianchangosaurus is the only known therizinosaur that possesses a tail with caudal centra that are oval in shape. [1]
Left cladogram is based on the phylogenetic analysis conducted by Pu et al. 2013, showing the relationships of Jianchangosaurus as a very primitive therizinosaur. [1] Right cladogram is based on Hartman et al. 2019 which has corroborated the position of Jianchangosaurus: [5]
|
|
The validity of Jianchangosaurus was called into question by Cau (2024), who considered it to be a junior synonym of the contemporary Beipiaosaurus since the diagnosis distinguishing the taxa was based on ontogenetically variable characters. [6]
Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.
Segnosaurus is a genus of therizinosaurid dinosaur that lived in what is now southeastern Mongolia during the Late Cretaceous, about 102–86 million years ago. Multiple incomplete but well-preserved specimens were discovered in the Gobi Desert in the 1970s, and in 1979 the genus and species Segnosaurus galbinensis were named. The generic name Segnosaurus means "slow lizard" and the specific name galbinensis refers to the Galbin region. The known material of this dinosaur includes the lower jaw, neck and tail vertebrae, the pelvis, shoulder girdle, and limb bones. Parts of the specimens have gone missing or become damaged since they were collected.
Beipiaosaurus is a genus of therizinosauroid theropod dinosaurs that lived in China during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the discovery of Yutyrannus, Beipiaosaurus were among the heaviest dinosaurs known from direct evidence to be feathered. Beipiaosaurus is known from three reported specimens. Numerous impressions of feather structures were preserved that allowed researchers to determine the feathering color which turned out to be brownish.
Alxasaurus is a genus of therizinosauroid theropod dinosaurs from the Early Cretaceous Bayin-Gobi Formation of Inner Mongolia. It is known from five specimens, recovered from the Bayin-Gobi in 1988, as part of the China-Canada Dinosaur Project. During their preparation, palaeontologists Dong Zhiming and Dale Russell noted strong similarities to Segnosaurus. In 1993, they described Alxasaurus and named its type species, A. elesitaiensis. While therizinosaurs had previously been tentatively seen as late-surviving basal sauropodomorphs, the description of Alxasaurus lent credence to the idea that they were instead highly derived coelurosaurs.
Huabeisaurus was a genus of dinosaur from the Late Cretaceous. It was a sauropod which lived in what is present-day northern China. The type species, Huabeisaurus allocotus, was first described by Pang Qiqing and Cheng Zhengwu in 2000. Huabeisaurus is known from numerous remains found in the 1990s, which include teeth, partial limbs and vertebrae. Due to its relative completeness, Huabeisaurus represents a significant taxon for understanding sauropod evolution in Asia. Huabeisaurus comes from Kangdailiang and Houyu, Zhaojiagou Town, Tianzhen County, Shanxi province, China. The holotype was found in the unnamed upper member of the Huiquanpu Formation, which is Late Cretaceous (?Cenomanian–?Campanian) in age based on ostracods, charophytes, and fission-track dating.
Neimongosaurus is a genus of herbivorous therizinosaur theropod dinosaur that lived in China during the Late Cretaceous period. Its fossils are known from the strata of the Iren Dabasu Formation. It is known from two specimens, discovered in 1999 by researchers from the Ministry of Land and Resources and described two years later. One species, N. yangi, is known, named after Chinese palaeontologist Yang Zhongjian.
Nanshiungosaurus is a genus of therizinosaurid that lived in what is now Asia during the Late Cretaceous of South China. The type species, Nanshiungosaurus brevispinus, was first discovered in 1974 and described in 1979 by Dong Zhiming. It is represented by a single specimen preserving most of the cervical and dorsal vertebrae with the pelvis. A supposed and unlikely second species, "Nanshiungosaurus" bohlini, was found in 1992 and described in 1997. It is also represented by vertebrae but this species however, differs in geological age and lacks authentic characteristics compared to the type, making its affinity to the genus unsupported.
Mahajangasuchus is an extinct genus of crocodyliform which had blunt, conical teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar. It was a fairly large predator, measuring up to 4 metres (13 ft) long.
Peloroplites is a monospecific genus of nodosaurid dinosaur from Utah that lived during the Late Cretaceous in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, Peloroplites cedrimontanus, is known from a partial skull and postcranial skeleton. It was named in 2008 by Kenneth Carpenter and colleagues. Peloroplites was 6 metres long and weighed 2 tonnes, making it one of the largest known nodosaurids, and came from a time when ankylosaurids and nodosaurids were attaining large sizes.
Polonosuchus is a genus of rauisuchid known from the late Triassic of Poland. It was a huge predator about 5–6 metres in length and, like all rauisuchians, was equipped with a large head of long sharp teeth. The legs were placed almost underneath the body, unlike most reptiles, which would have made it quite fast and a powerful runner. The appearance was very similar to that of the more known Postosuchus, of North America, and shared with the latter the ecological niche of the apex predator.
Fodonyx is an extinct genus of rhynchosaur from the middle Triassic epoch of Devon in England. Its fossils were discovered in Otter Sandstone Formation and were first assigned to Rhynchosaurus spenceri. This species was reassigned to its own genus, Fodonyx the holotype of which is EXEMS 60/1985/292, that described by David W. E. Hone and Michael J. Benton in 2008. In 2010, one skull was reassigned to the new genus Bentonyx. It is distinguished from other rhynchosaurs by a single autapomorphy, the ventral angling of the paraoccipital processes. In all other rhynchosaurs these processes angle dorsally or are horizontal. It is not known if this conferred any advantage to Fodonyx. Fodonyx was between 40 and 50 cm long.
Atacamatitan is a genus of titanosaurian sauropod dinosaurs from the Late Cretaceous Tolar Formation of Chile.
Manidens is an extinct genus of heterodontosaurid dinosaur from the Early Jurassic of Patagonia. It is a sister taxon of the closely related Pegomastax from South Africa. Fossils have been found in the Cañadón Asfalto Formation in Chubut Province, Argentina, considered to be originally dated to the Bajocian, latter were found to be from Toarcian beds.
Huehuecanauhtlus is a genus of hadrosauroid dinosaur known from the Late Cretaceous of Michoacán, western Mexico. It contains a single species, Huehuecanauhtlus tiquichensis.
Yurgovuchia is a genus of dromaeosaurid theropod dinosaurs that lived in North America during the Early Cretaceous period in what is now the Cedar Mountain Formation. It contains a single species, Yurgovuchia doellingi. The remains were discovered in Utah, United States.
Martharaptor is a genus of therizinosauroid theropod dinosaurs from the Early Cretaceous of the Cedar Mountain Formation in Utah. They can be distinguished from other therizinosauroids by means of several features of the skeleton which were intermediate between early therizinosaurs such as Falcarius and Beipiaosaurus, and more "advanced" members of the group like therizinosaurids. The deep and homogeneous hand claws clearly differ from the case in early therizinosauroids, but the foot has not yet acquired the robust morphology of therizinosaurids.
Europelta is a monospecific genus of nodosaurid dinosaur from Spain that lived during the Early Cretaceous in what is now the lower Escucha Formation of the Teruel Province. The type and only species, Europelta carbonensis, is known from two associated partial skeletons, and represents the most complete ankylosaur known from Europe. Europelta was named in 2013 by James I. Kirkland and colleagues. Europelta has an estimated length of 5 metres and weight of 1.3 tonnes, making it the largest member of the clade Struthiosaurini.
This Glossary explains technical terms commonly employed in the description of dinosaur body fossils. Besides dinosaur-specific terms, it covers terms with wider usage, when these are of central importance in the study of dinosaurs or when their discussion in the context of dinosaurs is beneficial. The glossary does not cover ichnological and bone histological terms, nor does it cover measurements.
Monquirasaurus is an extinct genus of giant short-necked pliosaurs who lived during the Early Cretaceous (Aptian) in what is now Colombia. One species is known, M. boyacensis, described in 2021 from an almost complete fossil skeleton, discovered in 1977 in the town of Villa de Leyva, located in Boyacá. Published descriptions of the holotype specimen estimate that it should reach a total size approaching 9–11 m (30–36 ft) in length and weighing 9–10 metric tons, making Monquirasaurus a large representative of the pliosaurids.
Baiyinosaurus is an extinct genus of stegosaurian dinosaurs from the Middle Jurassic Wangjiashan Formation of China. The genus contains a single species, B. baojiensis, known from a partial skeleton including cranial bones. The skeletal anatomy of Baiyinosaurus demonstrates transitional features between basal thyreophorans and stegosaurs. While many stegosaurs are known from China, Baiyinosaurus is the only one currently named from Gansu Province.
{{cite journal}}
: CS1 maint: DOI inactive as of November 2024 (link)