Unenlagiidae

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Unenlagiids
Temporal range: Late Cretaceous, 94–66  Ma
O
S
D
C
P
T
J
K
Pg
N
Possible Early Cretaceous record
Austroraptor Reconstruction.jpg
Life reconstruction of Austroraptor cabazai
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Paraves
Family: Unenlagiidae
Bonaparte, 1999
Type species
Unenlagia comahuensis
Novas & Puerta, 1997
Subfamilies
Synonyms
  • Unenlagiinia? Brum et al.,2021

Unenlagiidae is a proposed family of eumaniraptoran paravians that includes the subfamilies Unenlagiinae and possibly Halszkaraptorinae. Fossils of both subfamilies have been found in both Gondwanan and Laurasian deposits. The biology of the group suggests that some members were semiaquatic specialists. [1]

Classification

The family Unenlagiidae traditionally includes the same members as the previously named subfamily of Dromaeosauridae, Unenlagiinae, so Unenlagiidae was often seen as a synonym of Dromaeosauridae. However, since the 2010s, there have been subsequent studies that have questioned this placement, necessitating the revival of the family name. Some have placed unenlagiids as outside Dromaeosauridae, being the sister taxon or closely related to Avialae, [2] [3] while others have placed the newly recognized halszkaraptorines in the family, as basal deinonychosaurs outside Dromaeosauridae and Troodontidae. [4] Other recent studies supported the traditional placement of Halszkaraptorinae and Unenlagiinae in Dromaeosauridae, forming no distinct clade. [1] [5] In 2021, Brum et al. named the clade Unenlagiinia for a unified Unenlagia+Halszkaraptor clade, recovering them as basal dromaeosaurs. [6]

The following cladogram is from Motta et al., 2020, showing Unenlagiidae outside dromaeosaurids and more closely related to Avialae: [3]

Paraves

Troodontidae

Eumaniraptora

Dromaeosauridae

Microraptoria

Unenlagiidae

Austroraptor

Buitreraptor

Unenlagia

Overoraptor

Rahonavis

Avialae
Archaeopterygidae

Alcmonavis

Wellnhoferia

Archaeopteryx

Anchiornithidae

Xiaotingia

Anchiornis

Jeholornis

Pygostylia

Related Research Articles

<span class="mw-page-title-main">Troodontidae</span> Extinct family of bird-like dinosaurs

Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens, have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like Sinovenator, demonstrate striking anatomical similarities with Archaeopteryx and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves.

<span class="mw-page-title-main">Deinonychosauria</span> Extinct clade of dinosaurs

Deinonychosauria is a clade of paravian dinosaurs which lived from the Late Jurassic to the Late Cretaceous periods. Fossils have been found across the globe in North America, Europe, Africa, Asia, South America, and Antarctica, with fossilized teeth giving credence to the possibility that they inhabited Australia as well. This group of dinosaurs are known for their sickle-shaped toe claws and features in the shoulder bones.

<span class="mw-page-title-main">Dromaeosauridae</span> Family of theropod dinosaurs

Dromaeosauridae is a family of feathered coelurosaurian theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek δρομαῖος (dromaîos), meaning 'running at full speed', 'swift', and σαῦρος (saûros), meaning 'lizard'. In informal usage, they are often called raptors, a term popularized by the film Jurassic Park; several genera include the term "raptor" directly in their name, and popular culture has come to emphasize their bird-like appearance and speculated bird-like behavior.

<i>Neuquenraptor</i> Extinct genus of dinosaurs

Neuquenraptor is a genus of dromaeosaurid theropod dinosaurs that lived in South America during the Late Cretaceous in what is now the Portezuelo Formation of Argentina. It is one of the first dromaeosaurids found in the Southern Hemisphere.

<i>Buitreraptor</i> Dromaeosaurid dinosaur genus from the Late Cretaceous

Buitreraptor is a genus of dromaeosaurid dinosaurs that lived during the Late Cretaceous of Argentina at the Candeleros Formation. Buitreraptor was described in 2005 and the type species is Buitreraptor gonzalezorum. It was rooster-sized and had a very elongated head with many small teeth.

<i>Rahonavis</i> Extinct genus of dinosaurs

Rahonavis is a genus of bird-like theropods from the Late Cretaceous of what is now northwestern Madagascar. It is known from a partial skeleton found by Catherine Forster and colleagues in Maevarano Formation rocks at a quarry near Berivotra, Mahajanga Province. Rahonavis was a small predator, at about 70 centimetres (2.3 ft) long and 0.45-2.27 kg, with the typical Velociraptor-like raised sickle claw on the second toe. It was originally the first African coelurosaur until the Nqwebasaurus was discovered in 2000.

<i>Pyroraptor</i> Extinct genus of dinosaurs

Pyroraptor is an extinct genus of paravian dinosaur, probably a dromaeosaurid or unenlagiid, from the Late Cretaceous Ibero-Armorican island, of what is now southern France and northern Spain. It lived during the late Campanian and early Maastrichtian stages, approximately 83.5 to 70.6 million years ago. It is known from a single partial specimen that was found in Provence in 1992, after a forest fire. The animal was named Pyroraptor olympius by Allain and Taquet in 2000.

Unenlagia is a genus of dromaeosaurid theropod dinosaur that lived in South America during the Late Cretaceous period. The genus Unenlagia has been assigned two species: U. comahuensis, the type species described by Novas and Puerta in 1997, and U. paynemili, described by Calvo et al. in 2004.

<span class="mw-page-title-main">Scansoriopterygidae</span> Extinct family of dinosaurs

Scansoriopterygidae is an extinct family of climbing and gliding maniraptoran dinosaurs. Scansoriopterygids are known from five well-preserved fossils, representing four species, unearthed in the Tiaojishan Formation fossil beds of Liaoning and Hebei, China.

<i>Hulsanpes</i> Extinct genus of dinosaurs

Hulsanpes is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. Hulsanpes represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae.

<span class="mw-page-title-main">Archaeopterygidae</span> Family of dinosaurs

Archaeopterygidae is a group of maniraptoran dinosaurs, known from the latest Jurassic and earliest Cretaceous of Europe. In most current classifications, it contains only the genera Archaeopteryx and Wellnhoferia. As its name suggests, Protarchaeopteryx was also once referred to this group, but most paleontologists now consider it an oviraptorosaur. Other referred genera, like Jurapteryx, Wellnhoferia, and "Proornis", are probably synonymous with Archaeopteryx or do not belong into this group. Jinfengopteryx was originally described as an archaeopterygid, though it was later shown to be a troodontid. A few studies have recovered Anchiornis and Xiaotingia to also be members of the Archaeopterygidae, though most subsequent analyses have failed to arrive at the same result. Uncertainties still exist, however, and it may not be possible to confidently state whether archaeopterygids are more closely related to modern birds or to deinonychosaurs barring new and better specimens of relevant species. Teeth attributable to archaeopterygids are known from the earliest Cretaceous (Berriasian) Cherves-de-Cognac locality and the Angeac-Charente bonebed of France.

<span class="mw-page-title-main">Unenlagiinae</span> Extinct subfamily of dinosaurs

Unenlagiinae is a subfamily of long-snouted paravian theropods. They are traditionally considered to be members of Dromaeosauridae, though some authors place them into their own family, Unenlagiidae, alongside the subfamily Halszkaraptorinae. Unenlagiines are known from South America, with possible unenlagiines being known from North America, Madagascar, Europe, and even Australia.

<span class="mw-page-title-main">Avialae</span> Clade including all birds and their ancestors

Avialae is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used.

<span class="mw-page-title-main">Paraves</span> Clade of all dinosaurs that are more closely related to birds than to oviraptorosaurs

Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avialans, which include diverse extinct taxa as well as the over 10,000 species of living birds. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research.

<i>Epidexipteryx</i> Extinct genus of dinosaurs

Epidexipteryx is a genus of small paravian dinosaurs, known from one fossil specimen in the collection of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. Epidexipteryx represents the earliest known example of ornamental feathers in the fossil record.

<i>Austroraptor</i> Genus of theropod dinosaurs

Austroraptor is a genus of dromaeosaurid theropod dinosaur that lived during the Campanian and Maastrichtian ages of the Late Cretaceous period in what is now Argentina.

<span class="mw-page-title-main">Saurornitholestinae</span> Extinct subfamily of dinosaurs

Saurornitholestinae is a subfamily of the theropod group Dromaeosauridae. The saurornitholestines currently include three monotypic genera: Atrociraptor marshalli, Bambiraptor feinbergi, and Saurornitholestes langstoni. All are medium-sized dromaeosaurs from the Late Cretaceous of western North America. The group was originally recognized by Longrich and Currie as the sister taxon to a clade formed by the Dromaeosaurinae and Velociraptorinae. However, not all phylogenetic analyses recover this group and/or with the same proposed genera.

<i>Xiaotingia</i> Extinct genus of dinosaurs

Xiaotingia is a genus of anchiornithid theropod dinosaur from Middle Jurassic or early Late Jurassic deposits of western Liaoning, China, containing a single species, Xiaotingia zhengi.

<span class="mw-page-title-main">Anchiornithidae</span> Extinct family of reptiles

Anchiornithidae is a family of eumaniraptorans which could be the basalmost family of birds in the clade Avialae. Anchiornithids have been classified at varying positions in the maniraptoran tree, with some scientists classifying them as a distinct family, a basal subfamily of Troodontidae, members of Archaeopterygidae, or an assemblage of dinosaurs that are an evolutionary grade within Avialae or Paraves.

<i>Overoraptor</i> Extinct genus of theropod dinosaurs

Overoraptor is an extinct genus of paravian theropod of uncertain affinities from the Late Cretaceous Huincul Formation of Argentinian Patagonia. The genus contains a single species, O. chimentoi, known from several bones of the hands, feet, and hips alongside some vertebrae.

References

  1. 1 2 Cau, A.; Beyrand, V.; Voeten, D.; Fernandez, V.; Tafforeau, P.; Stein, K.; Barsbold, R.; Tsogtbaatar, K.; Currie, P.; Godrfroit, P. (6 December 2017). "Synchrotron scanning reveals amphibious ecomorphology in a new clade of bird-like dinosaurs". Nature . 552 (7685): 395–399. Bibcode:2017Natur.552..395C. doi:10.1038/nature24679. PMID   29211712. S2CID   4471941.
  2. Federico L. Agnolín and Fernando E. Novas (2013). Avian ancestors. A review of the phylogenetic relationships of the theropods Unenlagiidae, Microraptoria, Anchiornis and Scansoriopterygidae. SpringerBriefs in Earth System Sciences. pp. 1–96. doi:10.1007/978-94-007-5637-3. ISBN   978-94-007-5636-6. S2CID   199493087.
  3. 1 2 Matías J. Motta; Federico L. Agnolín; Federico Brissón Egli; Fernando E. Novas (2020). "New theropod dinosaur from the Upper Cretaceous of Patagonia sheds light on the paravian radiation in Gondwana". The Science of Nature. 107 (3): Article number 24. Bibcode:2020SciNa.107...24M. doi:10.1007/s00114-020-01682-1. hdl: 11336/135530 . PMID   32468191. S2CID   218913199.
  4. Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019). "A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight". PeerJ. 7: e7247. doi: 10.7717/peerj.7247 . PMC   6626525 . PMID   31333906.
  5. Cau, Andrea (2020-02-25). "The body plan of Halszkaraptor escuilliei (Dinosauria, Theropoda) is not a transitional form along the evolution of dromaeosaurid hypercarnivory". PeerJ. 8: e8672. doi: 10.7717/peerj.8672 . ISSN   2167-8359. PMC   7047864 . PMID   32140312.
  6. Brum, Arthur S.; Pêgas, Rodrigo V.; Bandeira, Kamila L.N.; Souza, Lucy G.; Campos, Diogenes A.; Kellner, Alexander W.A. (2021). "A new Unenlagiinae (Theropoda: Dromaeosauridae) from the Late Cretaceous of Brazil". Papers in Palaeontology. 7 (4): 2075–2099. doi:10.1002/spp2.1375. S2CID   238854675.