Unenlagiidae

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Unenlagiids
Temporal range: Late Cretaceous
Austroraptor Reconstruction.jpg
Life reconstruction of Austroraptor cabazai
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Paraves
Family: Unenlagiidae
Bonaparte, 1999
Type species
Unenlagia comahuensis
Novas & Puerta, 1997
Subfamilies
Synonyms
  • Unenlagiinia? Brum et al.,2021

Unenlagiidae is a proposed family of eumaniraptoran paravians that includes the subfamilies Unenlagiinae and possibly Halszkaraptorinae. Fossils of both subfamilies have been found in both Gondwanan and Laurasian deposits. The biology of the group suggests that some members were semiaquatic specialists. [1]

Classification

The family Unenlagiidae traditionally includes the same members as the previously named subfamily of Dromaeosauridae, Unenlagiinae, so Unenlagiidae was often seen as a synonym of Dromaeosauridae. However, since the 2010s, there have been subsequent studies that have questioned this placement, necessitating the revival of the family name. Some have placed unenlagiids as outside Dromaeosauridae, being the sister taxon or closely related to Avialae, [2] [3] while others have placed the newly recognized halszkaraptorines in the family, as basal deinonychosaurs outside Dromaeosauridae and Troodontidae. [4] Other recent studies supported the traditional placement of Halszkaraptorinae and Unenlagiinae in Dromaeosauridae, forming no distinct clade. [1] [5] In 2021, Brum et al. named the clade Unenlagiinia for a unified Unenlagia+Halszkaraptor clade, recovering them as basal dromaeosaurs. [6]

The following cladogram is from Motta et al., 2020, showing Unenlagiidae outside dromaeosaurids and more closely related to Avialae: [3]

Paraves

Related Research Articles

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<span class="mw-page-title-main">Dromaeosauridae</span> Family of theropod dinosaurs

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Unenlagia is a genus of dromaeosaurid theropod dinosaur that lived in South America during the Late Cretaceous period. The genus Unenlagia has been assigned two species: U. comahuensis, the type species described by Novas and Puerta in 1997, and U. paynemili, described by Calvo et al. in 2004.

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References

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  2. Federico L. Agnolín and Fernando E. Novas (2013). Avian ancestors. A review of the phylogenetic relationships of the theropods Unenlagiidae, Microraptoria, Anchiornis and Scansoriopterygidae. SpringerBriefs in Earth System Sciences. pp. 1–96. doi:10.1007/978-94-007-5637-3. ISBN   978-94-007-5636-6. S2CID   199493087.
  3. 1 2 Matías J. Motta; Federico L. Agnolín; Federico Brissón Egli; Fernando E. Novas (2020). "New theropod dinosaur from the Upper Cretaceous of Patagonia sheds light on the paravian radiation in Gondwana". The Science of Nature. 107 (3): Article number 24. Bibcode:2020SciNa.107...24M. doi:10.1007/s00114-020-01682-1. hdl: 11336/135530 . PMID   32468191. S2CID   218913199.
  4. Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019). "A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight". PeerJ. 7: e7247. doi: 10.7717/peerj.7247 . PMC   6626525 . PMID   31333906.
  5. Cau, Andrea (2020-02-25). "The body plan of Halszkaraptor escuilliei (Dinosauria, Theropoda) is not a transitional form along the evolution of dromaeosaurid hypercarnivory". PeerJ. 8: e8672. doi: 10.7717/peerj.8672 . ISSN   2167-8359. PMC   7047864 . PMID   32140312.
  6. Brum, Arthur S.; Pêgas, Rodrigo V.; Bandeira, Kamila L.N.; Souza, Lucy G.; Campos, Diogenes A.; Kellner, Alexander W.A. (2021). "A new Unenlagiinae (Theropoda: Dromaeosauridae) from the Late Cretaceous of Brazil". Papers in Palaeontology. 7 (4): 2075–2099. doi:10.1002/spp2.1375. S2CID   238854675.