Xixianykus Temporal range: Late Cretaceous, | |
---|---|
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Alvarezsauridae |
Genus: | † Xixianykus Xu et al, 2010 |
Species: | †X. zhangi |
Binomial name | |
†Xixianykus zhangi Xu et al., 2010 | |
Xixianykus is a genus of alvarezsaurid theropod dinosaur from the late Cretaceous period of China. [1] Known from a slab of rock which contains a partial postcranial skeleton unearthed in the Late Cretaceous (Coniacian-Santonian) Majiacun Formation of the Xixia Basin in Henan, China.
The fossil, holotype XMDFEC V0011, was found in the Majiacun Formation in Henan Province, China. The fossil consists of a partial skeleton, without the skull. Parts of the hind legs, pelvis and spine has also been recovered. Xixianykus is among the oldest of the derived alvarezsauroids, the parvicursorines, dated to the Santonian–Coniacian, as opposed to other parvicursorines, which are either Campanian or Maastrichtian. [2]
The type species is Xixianykus zhangi described in 2010 by Xu Xing. The genus name refers to Xixia County and to the Greek word for claw ("onyx"). The latter is a common element used in names for Alvarezsaurids. The species name is in honor of Zhang Xinglao.
Xixianykus was a small animal, about 50 centimetres (20 in) long and 20 centimetres (7.9 in) high, and may have weighed just about 0.74 kg. It is one of the smallest-known dinosaurs (not counting avians). It appears to have many adaptations towards a cursorial (running) lifestyle. It was about 50 centimetres long but had 20-centimetre-long (7.9 in) legs and a short femur combined with a long tibia and metatarsus, as well as including an arctometatarsalian pes, a highly modified structure in the feet of theropod dinosaurs used to enhance cursorial capability, which was found within the known fossil material, and these are all good indicators of it being a fast runner. [2] It was probably covered in feathers and had small, robust arms with one large claw for each hand, as well as a tubular snout used to feed likely on colonial insects.
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests.
Anserimimus is a genus of ornithomimid theropod dinosaur, from the Late Cretaceous Period of what is now Mongolia. It was a lanky, fast-running animal, possibly an omnivore. From what fossils are known, it probably closely resembled other ornithomimids, except for its more powerful forelimbs.
Yixianosaurus is a maniraptoran theropod dinosaur genus from the Early Cretaceous of China.
Mononykus is a genus of alvarezsaurid dinosaur that lived during the Late Cretaceous in what is now Asia on the Nemegt Formation, about 70 million years ago.
Sinusonasus is a genus of dinosaurs from the Early Cretaceous Period, recovered from the Yixian Formation. It lived in what is now the Liaoning Province of China. Sinusonasus was a theropod, specifically a troodontid dinosaur.
Patagonykus is a genus of theropod dinosaur from the Upper Cretaceous of Argentina. This alvarezsauroid was discovered in exposures of the Portezuelo Formation (Turonian-Coniacian) of the Rio Neuquén Subgroup in the Neuquén Basin, Neuquen Province of Patagonia, Argentina. The holotype consists of an incomplete but well-preserved skeleton, lacking a skull, but including many vertebrae, the coracoids, a partial forelimb, pelvic girdle, and hindlimbs. Patagonykus has been classed with the Alvarezsauridae, a family which includes such taxa as the Mongolian Mononykus and the Argentinian Alvarezsaurus. In 2010 Gregory S. Paul estimated its length at 1 meter and its weight at 3.5 kg.
Graciliraptor is a genus of theropod dinosaur from the early Cretaceous Period. It is a microraptorine dromaeosaurid.
Achillesaurus is a genus of alvarezsaurid theropod dinosaur from the Santonian-age Upper Cretaceous Bajo de la Carpa Formation of Rio Negro, Argentina. It was named in reference to Achilles' heel, because diagnostic features are found there for these animals. The genus was a relatively large, basal alvarezsaurid, and a contemporary of Alvarezsaurus. Achillesaurus is based on MACN-PV-RN 1116, a partial skeleton including a sacral vertebra, four tail vertebrae, part of the left thighbone, shin and foot, and the left ilium. Agustín Martinelli and Ezequiel Vera, who described the specimen, performed a phylogenetic analysis and found their new genus to be an alvarezsaurid with an unresolved relationship to Alvarezsaurus and more derived alvarezsaurids. Makovicky, Apesteguía & Gianechini (2012) argued that Achillesaurus might actually be a junior synonym of Alvarezsaurus which, according to the authors, "is known from the same formation and from which it [i.e. Achillesaurus] differs trivially."
Luanchuanraptor is a genus of dromaeosaurid theropod dinosaurs from the Late Cretaceous of China. The genus is based on a partial skeleton from the Qiupa Formation in Luanchuan, Henan. They were medium-sized dromaeosaurids, the first Asian dromaeosaurid taxa described from outside the Gobi Desert or northeastern China.
Albertonykus is an alvarezsaurid dinosaur from the Maastrichtian-age rocks of the Horseshoe Canyon Formation of Alberta, Canada. It is known from forelimb and hindlimb remains from multiple individuals. All but two of the specimens come from a bonebed dominated by Albertosaurus, located at the top of Unit 4 of the Horseshoe Canyon Formation, dating to ~68.5 million years ago.
Alvarezsauroidea is a group of small maniraptoran dinosaurs. Alvarezsauroidea, Alvarezsauridae, and Alvarezsauria are named for the historian Gregorio Álvarez, not the more familiar physicist Luis Alvarez, or his son geologist Walter Alvarez who jointly proposed that the Cretaceous–Paleogene extinction event was caused by an impact event. The group was first formally proposed by Choiniere and colleagues in 2010, to contain the family Alvarezsauridae and non-alvarezsaurid alvarezsauroids, such as Haplocheirus, which is the basalmost of the Alvarezsauroidea. The discovery of Haplocheirus extended the stratigraphic evidence for the group Alvarezsauroidea about 63 million years further in the past. The division of Alvarezsauroidea into the Alvarezsauridae and the non-alvarezsaurid alvarezsauroids is based on differences in their morphology, especially in their hand morphology.
Xixiasaurus is a genus of troodontid dinosaur that lived during the Late Cretaceous Period in what is now China. The only known specimen was discovered in Xixia County, Henan Province, in central China, and became the holotype of the new genus and species Xixiasaurus henanensis in 2010. The names refer to the areas of discovery, and can be translated as "Henan Xixia lizard". The specimen consists of an almost complete skull, part of the lower jaw, and teeth, as well as a partial right forelimb.
The Majiacun Formation is a Santonian to Coniacian geologic formation in China. Dinosaur remains are among the fossils that have been recovered from the formation.
Qiupalong is an extinct genus of ornithomimosaurian theropod that was discovered in the Late Cretaceous Qiupa Formation of Henan, China. The genus contains a single species, Q. henanensis, the specific epithet for which was named for the province of Henan. Uniquely, Qiupalong is one of the few Late Cretaceous non-avian dinosaurs known from both Asia and Laramidia. Specimens from Russia and Alberta have been referred to the genus without being assigned to the type species.
Macroelongatoolithus is an oogenus of large theropod dinosaur eggs, representing the eggs of giant caenagnathid oviraptorosaurs. They are known from Asia and from North America. Historically, several oospecies have been assigned to Macroelongatoolithus, however they are all now considered to be a single oospecies: M. carlylensis.
Linhenykus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous of Inner Mongolia, China. It is the most basal known member of the Parvicursorinae. The genus gets its name from Linhe, a city near the site where the fossil was first found and Greek nykus, "claw". The specific name is derived from Greek monos, "single", and daktylos, "finger", a reference to the fact that it is the only known non-avian dinosaur to have had but a single digit.
Aorun is a genus of carnivorous theropod dinosaur first discovered in 2006, with its scientific description published in 2013. It is possibly one of the oldest known coelurosaurian dinosaurs and is estimated to have lived ~161.6 million years ago during the Late Jurassic Period.
Zhanghenglong is an extinct genus of herbivorous hadrosauroid iguanodont dinosaur known from the Late Cretaceous Majiacun Formation in Xixia County of Henan Province, China. It contains a single species, Zhanghenglong yangchengensis, represented by a disarticulated and partial cranium and postcranial skeleton.
Qiupanykus is a genus of alvarezsaurid coelurosaur theropod from the Late Cretaceous Qiupa Formation of southern China.
Fujianvenator is an extinct genus of anchiornithid theropod dinosaur from the Late Jurassic Nanyuan Formation of Fujian Province, China. The genus contains a single species, F. prodigiosus, known from a partial articulated skeleton. It is suggested to possibly have had a terrestrial or wading lifestyle.