Zhongjianornis Temporal range: Early Cretaceous, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Clade: | Euornithes |
Genus: | † Zhongjianornis Zhou, Zhang & Li, 2010 |
Species: | †Z. yangi |
Binomial name | |
†Zhongjianornis yangi Zhou, Zhang & Li, 2010 | |
Zhongjianornis is a genus of beaked, pigeon-sized birds from the early Cretaceous period of China. It is known from one fossil found at Jianchang, Liaoning Province, in rocks of the Jiufotang Formation, representing the type species Zhongjianornis yangi.
The holotype specimen is in the collection of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing, and the species Z. yangi is named for the IVPP's founder, Yang Zhongjian. [1] This specimen is catalogued under the accession number IVPP V15900. It consists of a complete skeleton, possibly only missing a few tail vertebrae.
Z. yangi was initially believed to be the most primitive (though not earliest) bird species that completely lacked teeth, and one of the most basal birds known. It was described by Zhou, Zhang and Li, in a paper published in January 2010 (though the paper had appeared on the Internet during 2009). [2] They found that Zhongjianornis was more primitive than Confuciusornis , and that it predated the split between euornithean and enantiornithean birds. The authors also reported that Zhongjianornis shared features with very basal birds like Jeholornis , Sapeornis and Confuciusornis , including a highly expanded deltopectoral crest, and suggest that this was an early flight architecture that was replaced by an enlarged, keeled breastbone in more advanced birds.
However, later analysis of the anatomy of Z. yangi cast doubt on this interpretation of the bird as very primitive. Research published in 2012 found that some of the apparently primitive features were due to poor preservation of the bones, and that Zhongjianornis is more similar in anatomy to primitive euornitheans like Chaoyangia and Schizooura . A phylogenetic analysis showed that a very primitive position in the bird family tree for Z. yangi is unlikely given current evidence. [3]
Caudipteryx is a genus of small oviraptorosaur dinosaurs that lived in Asia during the Early Cretaceous, around 124.6 million years ago. They were feathered and extremely birdlike in their overall appearance, to the point that some paleontologists suggested it was a bird. Two species have been described: C. zoui, in 1998, and C. dongi, in 2000.
Confuciusornis is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, Confuciusornis had a toothless beak, but closer and later relatives of modern birds such as Hesperornis and Ichthyornis were toothed, indicating that the loss of teeth occurred convergently in Confuciusornis and living birds. It was thought to be the oldest known bird to have a beak, though this title now belongs to an earlier relative Eoconfuciusornis. It was named after the Chinese moral philosopher Confucius. Confuciusornis is one of the most abundant vertebrates found in the Yixian Formation, and several hundred complete specimens have been found.
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Beipiaosaurus is a genus of therizinosauroid theropod dinosaurs that lived in Asia during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the discovery of Yutyrannus, Beipiaosaurus were among the heaviest dinosaurs known from direct evidence to be feathered. Beipiaosaurus is known from three reported specimens. Numerous impressions of feather structures were preserved that allowed researchers to determine the feathering color which turned out to be brownish.
Yanornis is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.
Jeholornis is a genus of avialan dinosaurs that lived between approximately 122 and 120 million years ago during the early Cretaceous Period in China. Fossil Jeholornis were first discovered in the Jiufotang Formation in Hebei Province, China and additional specimens have been found in the older Yixian Formation.
Gansus is a genus of aquatic birds that lived during the Aptian age of the Early Cretaceous (Aptian-Albian) period in what are now Gansu and Liaoning provinces, western China. The rock layers from which their fossils have been recovered are dated to 120 million years ago. It was first described in 1984 on the basis of an isolated left leg. It is the oldest-known member of the Ornithurae, the group which includes modern birds (Neornithes) and extinct related groups, such as Ichthyornis and Hesperornithes.
Sapeornis is a monotypic genus of avialan dinosaurs which lived during the early Cretaceous period. Sapeornis contains only one species, Sapeornis chaoyangensis.
Sinopterus is a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. Historically, there were multiple species attributed to the genus although only one is considered to be valid. Sinopterus is known for its proportionally large skull, which has a birdlike pointed beak, a long bony crest that starts with a tall premaxilla and goes back along the middle of the skull to form a point overhanging the rear of the skull, and its lack of teeth.
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their closest extinct relatives. Confuciusornithidae contains four genera, possessing both shafted and non-shafted (downy) feathers. They are also noted for their distinctive pair of ribbon-like tail feathers of disputed function.
Cathayornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known definitively from only one species, Cathayornis yandica, one of the first Enantiornithes found in China. Several additional species were once incorrectly classified as Cathayornis, and have since been reclassified or regarded as nomina dubia.
Yixianornis is a bird genus from the early Cretaceous period. Its remains have been found in the Jiufotang Formation at Chaoyang dated to the early Aptian age, around 120 million years ago. Only one species, Yixianornis grabaui, is known at present. The specific name, grabaui, is named after American paleontologist Amadeus William Grabau, who surveyed China in the early 20th century.
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces.
Protopteryx is an extinct bird and possibly the basalmost enantiornithean, from the Cretaceous period. The type species is P. fengningensis. It was first discovered in the Sichakou Member of the Yixian Formation or Huajiying Formation of Hebei Province, northern China, dating from 131 Ma ago. Protopteryx has been found in the Daibeigou formation, as well. The name Protopteryx means "primitive feather": "proto-" meaning "the first of" and "-pteryx" meaning "feather" or "wing." The name comes from the fact that Protopteryx feathers are more primitive than those of modern birds, such as the two elongated tail feathers that lack barbs and rami.
Zhongornis is a genus of primitive maniraptoran dinosaurs that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008.
Eoconfuciusornis a genus of extinct avialan that lived 131 Ma ago, in the Early Cretaceous of China. It is the oldest known bird to have a beak.
Chaoyangia is an extinct genus of euornithean birds, containing the single species Chaoyangia beishanensis. This species is known from a single fossil specimen consisting of a partial skeleton including vertebra, ribs, hips, and upper legs. The specimen was discovered in the Jiufotang Formation near the city of Chaoyang in Liaoning province, China. This rock formation has been dated to the Aptian age of the Early Cretaceous period, 120 million years ago.
Songlingornithidae is a family of basal euornithean dinosaurs from the Early Cretaceous of China. All known specimens come from the Jiufotang Formation and the Yixian Formation, dating to the early Barremian and Aptian ages, 125–120 million years ago.
Euornithes is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to Sinornis.
Bohaiornithidae is a group of early predatory enantiornitheans from the early Cretaceous Period of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125–120 million years ago. Bohaiornithidae was first coined as a family of enantiornithean birds by Wang and colleagues in 2014. They defined it as the natural group formed by all descendants of the common ancestor of the type species, Bohaiornis guoi, and Shenqiornis mengi.