Brodavis Temporal range: Late Cretaceous, | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Clade: | † Hesperornithes |
Family: | † Brodavidae Martin et al., 2012 |
Genus: | † Brodavis Martin et al., 2012 |
Type species | |
†Brodavis americanus Martin et al., 2012 | |
Species | |
|
Brodavis is a genus of freshwater hesperornithiform birds known from the Late Cretaceous (possibly Campanian and Maastrichtian stage) of North America and Asia. It was first described and named by Larry D. Martin, Evgeny N. Kurochkin and Tim T. Tokaryk in 2012 and assigned to a new monogeneric family, Brodavidae. Four species were described and assigned to Brodavis.
The type species, B. americanus, is known from the holotype left metatarsal, RSM P 2315.1 which was collected in the Maastrichtian-age Frenchman Formation of Canada.
B. baileyi is known from the holotype left metatarsal, UNSM 50665, which was collected in the Maastrichtian-age Hell Creek Formation of South Dakota, United States (dated to between 66.8 and 66 Ma ago [1] )
B. mongoliensis is known from the holotype left metatarsal, PIN 4491-8, which was collected in the Maastrichtian-age Nemegt Formation of Mongolia.
B. varneri was originally named as a second species of Baptornis by James Martin and Amanda Cordes-Person in 2007. It is based on the holotype left tarsometatarsal SDSM 68430 which was collected in the Campanian-age Sharon Springs Formation, lower Pierre Shale of southwestern South Dakota (dated to between 81.5 and 80.5 million years ago. [2] ). "Baptornis" varneri might represent a fourth species of Brodavis or belong to a separate genus.
Brodavis is considered to be the first freshwater and latest record of the order Hesperornithiformes. Although hesperornithiforms probably lost their flight abilities by the end of the Early Cretaceous, minimal pachyostosis in Brodavis suggests the possibility of some volant abilities. [3]
The Late Cretaceous is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after the white limestone known as chalk, which occurs widely in northern France and is seen in the white cliffs of south-eastern England, and which dates from this time.
Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 5 m (16 ft) from head to tail and 2,200 kg (4,900 lb) in body mass. It is known only from the Dakota Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Microstega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle.
Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as Hesperornis, Parahesperornis, Baptornis, Enaliornis, and Potamornis, all strong-swimming, predatory divers. Many of the species most specialized for swimming were completely flightless. The largest known hesperornithean, Canadaga arctica, may have reached a maximum adult length of 2.2 metres (7.2 ft).
Hesperornis is a genus of cormorant-like bird that spanned the first half of the Campanian age of the Late Cretaceous period. One of the lesser-known discoveries of the paleontologist O. C. Marsh in the late 19th century Bone Wars, it was an early find in the history of avian paleontology. Locations for Hesperornis fossils include the Late Cretaceous marine limestones from Kansas and the marine shales from Canada. Nine species are recognised, eight of which have been recovered from rocks in North America and one from Russia.
The Lance (Creek) Formation is a division of Late Cretaceous rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas. The Lance Formation is Late Maastrichtian in age, and shares much fauna with the Hell Creek Formation of Montana and North Dakota, the Frenchman Formation of southwest Saskatchewan, and the lower part of the Scollard Formation of Alberta.
Sphaerotholus is a genus of pachycephalosaurid dinosaur from the Upper Cretaceous of the western United States and Canada. To date, three species have been described: the type species, S. goodwini, from the Den-na-zin Member of the Kirtland Formation of San Juan County, New Mexico, USA; S. buchholtzae, from the Hell Creek Formation of western Carter County, Montana, USA and the Frenchman Formation of Saskatchewan, Canada; and S. edmontonensis, from the Horseshoe Canyon Formation of Alberta, Canada.
Hulsanpes is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. Hulsanpes represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae.
Ichthyornithes is an extinct group of toothed avialans very closely related to the common ancestor of all modern birds. They are known from fossil remains found throughout the late Cretaceous period of North America, though only one species, Ichthyornis dispar, is represented by complete enough fossils to have been named. Ichthyornitheans became extinct at the Cretaceous–Paleogene boundary, along with enantiornitheans, all other non-avian dinosaurs, and many other animal and plant groups.
Baptornis is a genus of flightless, aquatic birds from the Late Cretaceous, some 87-80 million years ago. The fossils of Baptornis advenus, the type species, were discovered in Kansas, which at its time was mostly covered by the Western Interior Seaway, a shallow shelf sea. It is now known to have also occurred in today's Sweden, where the Turgai Strait joined the ancient North Sea; possibly, it occurred in the entire Holarctic.
Alexornis is a genus of enantiornithine birds from the Bocana Roja Formation of Baja California, Mexico. This geological formation has been dated to the late Cretaceous period, and more specifically to the Campanian age, about 73 mya. The type and only known species is Alexornis antecedens. Its name means "Alex's ancestral bird"; Alexornis from the given name of ornithologist Alexander Wetmore + Ancient Greek ornis, "bird", and antecedens, Latin for "going before" or "ancestral".
Dolichorhynchops is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing three species, D. osborni, D. bonneri and D. tropicensis, as well as a questionably referred fourth species, D. herschelensis. Dolichorhynchops was an oceangoing prehistoric reptile. Its Greek generic name means "long-nosed face".
Cimolopteryx is a prehistoric bird genus from the Late Cretaceous Period. It is currently thought to contain only a single species, Cimolopteryx rara. The only specimen confidently attributed to C. rara was found in the Lance Formation of Wyoming, dating to the end of the Maastrichtian age, which ended about 66 million years ago. The dubious species "Cimolopteryx" maxima has been described from both the Lance Formation and the Hell Creek Formation of Montana. The humeral end of a left coracoid from the Frenchman Formation of southern Saskatchewan has also been attributed to the genus.
Teviornis is a genus of extinct birds. One species has been described, T. gobiensis. It lived in the Maastrichtian stage at the end of the Late Cretaceous period, some 70 million years ago. It is known from fossils collected from the Nemegt Formation of Gobi, south Mongolia.
Parahesperornis is a genus of prehistoric flightless birds from the Late Cretaceous. Its range in space and time may have been extensive, but its remains are rather few and far between, at least compared with its contemporary relatives in Hesperornis. Remains are known from central North America, namely the former shallows of the Western Interior Seaway in Kansas. Found only in the upper Niobrara Chalk, these are from around the Coniacian-Santonian boundary, 85-82 million years ago (mya).
Titanoceratops is a controversial genus of herbivorous ceratopsian dinosaur. It was a giant chasmosaurine ceratopsian that lived in the Late Cretaceous period in what is now New Mexico. Titanoceratops was named for its large size, being one of the largest known horned dinosaurs and the type species was named T. ouranos, after Uranus (Ouranos), the father of the Greek titans. It was named in 2011 by Nicholas R. Longrich for a specimen previously referred to Pentaceratops. Longrich believed that unique features found in the skull reveal it to have been a close relative of Triceratops, classified within the subgroup Triceratopsini. However, other researchers have expressed skepticism, and believe "Titanoceratops" to simply be an unusually large, old specimen of Pentaceratops.
Acheroraptor is an extinct genus of dromaeosaurid theropod dinosaur known from the latest Maastrichtian Hell Creek Formation of Montana, United States. It contains a single species, Acheroraptor temertyorum. A. temertyorum is one of the two geologically youngest known species of dromaeosaurids, the other being Dakotaraptor, which is also known from Hell Creek. A basal cousin of Velociraptor, Acheroraptor is known from upper and lower jaw material.
Tototlmimus is an extinct genus of ornithomimid dinosaur. Its remains were found from the late Cretaceous Packard Formation, in the Sonora state, México.
Fumicollis is a genus of prehistoric flightless birds from the Late Cretaceous (Coniacian-Santonian) Niobrara Chalk of Kansas.
Chupkaornis is a genus of prehistoric flightless birds from the Late Cretaceous (Coniacian-Santonian) Kashima Formation of Hokkaido, Japan.
Maip is a genus of large megaraptorid theropod dinosaur from the Late Cretaceous (Maastrichtian) Chorrillo Formation of Santa Cruz, Argentina. The genus contains a single species, M. macrothorax, known from an incomplete, disarticulated skeleton. Maip may represent the largest megaraptorid known from South America, and possibly the world.