Alexornis Temporal range: Late Cretaceous, | |
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Life reconstruction of Alexornis antecedens | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Clade: | † Enantiornithes |
Order: | † Alexornithiformes Brodkorb, 1976 |
Family: | † Alexornithidae Brodkorb, 1976 |
Genus: | † Alexornis Brodkorb, 1976 |
Species: | †A. antecedens |
Binomial name | |
†Alexornis antecedens Brodkorb, 1976 | |
Alexornis is a genus of enantiornithine birds from the Bocana Roja Formation of Baja California, Mexico. [1] This geological formation has been dated to the late Cretaceous period, and more specifically to the Cenomanian To Turonian age, about 93.6 mya. The type and only known species is Alexornis antecedens. Its name means "Alex's ancestral bird"; Alexornis from the given name of ornithologist Alexander Wetmore + Ancient Greek ornis, "bird", and antecedens, Latin for "going before" or "ancestral". [2]
A. antecedens is known only from a single fragmentary skeleton including shoulder, wing, and leg bones, but lacking a skull. It was about the size of a sparrow, weighing only about 35.6 grams (1.26 oz). [3] [4] The specimen was discovered in 1971 and described and named in 1976 by Pierce Brodkorb. Brodkorb considered the species to be similar to some modern birds, and at the time, recognized it as one of the few known Cretaceous "land birds" after Gobipteryx minuta (most other Cretaceous birds at the time were thought to be aquatic or semi-aquatic). [2]
Alexornis was originally described as an early member of the modern bird group and, more specifically, as a possible ancestor of the orders Coraciiformes (kingfishers and relatives) and Piciformes (woodpeckers and relatives). However, when the first recognized enantiornithine fossils were discovered, it soon became clear that Alexornis was an additional member of this group. [3] Enantiornithes are characterized by a reversed articulation of the scapula and coracoid bones in the shoulder, and in 1983 Larry Martin showed that Brodkorb has accidentally switched these two bones in his original description, confused by their reversed anatomy. [5]
In the past, Alexornis has been allied with the similarly sized Central Asian genus Kizylkumavis , based on characteristics which are known to unite them with other Cretaceous birds in the group Enantornithes. [3] Evgeny Kurochkin argued in 1996 that Alexornis formed a natural group, the Alexornithidae, with Kizylkumavis and a second type of central Asian bird, Sazavis . Among the enantiornithes, Alexornis shares many similarities of its skeletal anatomy with South American birds from about the same time period, including Martinavis and Elbretornis , and is united with them in the group Euenantiornithes. [6]
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Iberomesornis is a monotypic genus of enantiornithine bird of the Cretaceous of Spain.
Yanornis is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.
Ichthyornithes is an extinct group of toothed avialan dinosaurs very closely related to the common ancestor of all modern birds. They are known from fossil remains found throughout the late Cretaceous period of North America, though only two genera, Ichthyornis and Janavis, are represented by complete enough fossils to have been named. Ichthyornitheans became extinct at the Cretaceous–Paleogene boundary, along with enantiornitheans, all other non-avian dinosaurs, and many other animal and plant groups.
Nanantius is a genus of extinct enantiornithean avialan known from the Early Cretaceous of Australia.
Enantiornis is a genus of Enantiornithes. The type and only currently accepted species E. leali is from the Late Cretaceous Lecho Formation at El Brete, Argentina. It was described from specimen PVL-4035, a coracoid, proximal scapula and proximal humerus found close to each other and suspected to represent the left shoulder of a single individual.
Yungavolucris is a genus of enantiornitheans. It contains the single species Yungavolucris brevipedalis, which lived in the Late Cretaceous. The fossil bones were found in the Lecho Formation at estancia El Brete, Argentina."Yungavolucris brevipedalis" means "Short-footed Yungas bird". The generic name, Yungavolucris is after the Yungas region + the Latin volucris, which translates to "bird". The specific name brevipedalis is from the Latin brevis, which means "short", + pedalis, from the Latin pes, meaning "foot".
Lectavis is a genus of enantiornithine avialan. Their fossil bones have been recovered from the Late Cretaceous Lecho Formation at estancia El Brete, Argentina. The genus contains a single species, Lectavis bretincola.
Eoalulavis is a monotypic genus of enantiornithean bird that lived during the Barremian, in the Lower Cretaceous around 125 million years ago. The only known species is Eoalulavis hoyasi.
Concornis is a genus of enantiornithean birds which lived during the early Cretaceous period, in the late Barremian age about 125 million years ago. Its remains are known from the Calizas de La Huérgina Formation at Las Hoyas, Cuenca province, Spain. The single known species, Concornis lacustris, was described from the remains of one fairly complete individual skeleton.
Gurilynia is a genus of enantiornithine birds. One species is known, G. nessovi. It lived during the Maastrichtian stage of the Late Cretaceous period, between 70 and 66 mya. Gurilynia is known from fragmentary fossils found at the Gurilyn Tsav locality of the Nemegt Formation in south Gobi, Mongolia.
Apsaravis is a Mesozoic avialan genus from the Late Cretaceous. The single known species, Apsaravis ukhaana, lived about 78 million years ago, in the Campanian age of the Cretaceous period. Its fossilized remains were found in the Camel's Humps sublocality of the Djadokhta Formation, at Ukhaa Tolgod in the Gobi Desert of Mongolia. They were collected in the 1998 field season by the Mongolian Academy of Sciences/American Museum of Natural History Paleontological Expeditions. It was described by Norell and Clarke (2001).
Ornithothoraces is a group of avialan dinosaurs that includes all enantiornithes and the euornithes, which includes modern birds and their closest ancestors. The name Ornithothoraces means "bird thoraxes". This refers to the modern, highly advanced anatomy of the thorax that gave the ornithothoracines superior flight capability compared with more primitive avialans. This anatomy includes a large, keeled breastbone, elongated coracoids and a modified glenoid joint in the shoulder, and a semi-rigid rib cage. In spite of this at least the sternum seems to have developed convergently rather than being a true homology.
Avisauridae is a family of extinct enantiornithine dinosaurs from the Cretaceous period, distinguished by several features of their ankle bones. Depending on the definition used, Avisauridae is either a broad and widespread group of advanced enantiornithines, or a small family within that group, restricted to species from the Late Cretaceous of North and South America.
Soroavisaurus is a genus of enantiornithean birds related to Avisaurus. It lived during the Late Cretaceous of Argentina. The only known species, S. australis, is known from fossils collected from the Lecho Formation of Estancia El Brete, in the southern tip of the province of Salta, Argentina. A binominal name of this animal means "Southern sister Avisaur".
Elbretornis is an extinct genus of enantiornithine which existed in what is now Salta Province, Argentina during the late Cretaceous period.
Martinavis is a genus of enantiornithine birds which existed in what is now southern France, North America and Salta Province, Argentina during the late Cretaceous period. It was named by Cyril A. Walker, Eric Buffetaut and Gareth J. Dyke in 2007, and the type species is Martinavis cruzyensis.
Pengornithidae is a group of early enantiornithines from the early Cretaceous Period of China, with the putative member Falcatakely possibly extending this clade's range into the Late Cretaceous of Madagascar, and several putative pengornithids also hail from this formation. Specimens of these animals have been found both in the Huajiying Formation and Jiufotang Formation of Liaoning and Hebei provinces, dating from the Hauterivian age to the Aptian age.
Dunhuangia is a bird genus, belonging to the Enantiornithes, which during the Early Cretaceous lived in the area of present China.