Confuciusornis is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, Confuciusornis had a toothless beak, but closer and later relatives of modern birds such as Hesperornis and Ichthyornis were toothed, indicating that the loss of teeth occurred convergently in Confuciusornis and living birds. It was thought to be the oldest known bird to have a beak, though this title now belongs to an earlier relative Eoconfuciusornis. It was named after the Chinese moral philosopher Confucius. Confuciusornis is one of the most abundant vertebrates found in the Yixian Formation, and several hundred complete specimens have been found.
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
The Maevarano Formation is a Late Cretaceous sedimentary rock formation found in the Mahajanga Province of northwestern Madagascar. It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rivers that had greatly varying discharges. Notable animal fossils recovered include the theropod dinosaur Majungasaurus, the early bird Vorona, the paravian Rahonavis, the titanosaurian sauropod Rapetosaurus, and the giant frog Beelzebufo.
Yixianornis is a bird genus from the early Cretaceous period. Its remains have been found in the Jiufotang Formation at Chaoyang dated to the early Aptian age, around 120 million years ago. Only one species, Yixianornis grabaui, is known at present. The specific name, grabaui, is named after American paleontologist Amadeus William Grabau, who surveyed China in the early 20th century.
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces.
Protopteryx is an extinct bird and the most basal enantiornithean, from the Cretaceous period. The type species is P. fengningensis. It was first discovered in the Sichakou Member of the Yixian Formation or Huajiying Formation of Hebei Province, northern China, dating from 131 Ma ago. Protopteryx has been found in the Daibeigou formation, as well. The name Protopteryx means "primitive feather": "proto-" meaning "the first of" and "-pteryx" meaning "feather" or "wing." The name comes from the fact that Protopteryx feathers are more primitive than those of modern birds, such as the two elongated tail feathers that lack barbs and rami.
Zhongornis is a genus of primitive avialan that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008.
Shanweiniao is a genus of long-snouted enantiornithean birds from Early Cretaceous China. One species is known, Shanweiniao cooperorum. There is one known fossil, a slab and counterslab. The fossil is in the collection of the Dalian Natural History Museum, and has accession number DNHM D1878/1 and DNHM1878/2. It was collected from the Lower Cretaceous Dawangzhengzi Beds, middle Yixian Formation, from Lingyuan in the Liaoning Province, China.
Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.
Bohaiornis is a genus of enantiornithean birds. Fossils have been found from the Lower Cretaceous Jiufotang Formation of western Liaoning, China. The only known species, Bohaiornis guoi, was named by Dongyu Hu, Li Li, Lianhaim Hou and Xing Xu in 2011 on the basis of a fully articulated and well-preserved skeleton of a sub-adult. This specimen, LPM B00167, preserved two long, ribbon-like feathers attached to the tail rather than a fan of shorter pennaceous feathers. It was similar to the slightly older Eoenantiornis, but much larger in size. Bohaiornis is the type species of Bohaiornithidae, a family of large predatory enantiornitheans from the Early Cretaceous.
Euornithes is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to Sinornis.
Parabohaiornis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Parabohaiornis martini.
Bohaiornithidae is a group of early predatory enantiornitheans from the early Cretaceous Period of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125–120 million years ago. Bohaiornithidae was first coined as a family of enantiornithean birds by Wang and colleagues in 2014. They defined it as the natural group formed by all descendants of the common ancestor of the type species, Bohaiornis guoi, and Shenqiornis mengi.
Longusunguis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Longusunguis kurochkini.
Cruralispennia is an extinct genus of enantiornithean bird. The only known specimen of Cruralispennia was discovered in the Early Cretaceous Huajiying Formation of China and formally described in 2017. The type species of Cruralispennia is Cruralispennia multidonta. The generic name is Latin for "shin feather", while the specific name means "many-toothed". The holotype of Cruralispennia is IVPP 21711, a semi-articulated partial skeleton surrounded by the remains of carbonized feathers.
Parapengornis is an extinct genus of enantiornithine bird from the Lower Cretaceous of what is now China. The holotype specimen was discovered in the Jiufotang Formation near Lingyuan, western Liaoning province, and was catalogued as IVPP V18687. The nearly complete, articulated specimen is preserved on a slab and has impressions of pennaceous feathers. Only parts of the sternum, the left hand, and right foot are missing. In 2015, it became the basis of the new genus and species Parapengornis eurycaudatus, named by the Chinese palaeontologists Han Hu, Jingmai K. O’Connor, and Zhonghe Zhou. The generic name consists of the Latin word para and the name of the related genus Pengornis, indicating their close relationship. The name Pengornis is itself derived from "Peng", a mythological bird from Chinese folklore, and ornis, which means bird in Greek. The specific name is derived from the Latin words eury, meaning broad, and caudatus, meaning tail, in reference to the broad and expanded pygostyle. A nearly complete specimen formerly assigned to Pengornis was also reassigned to Parapengornis by these authors.
Chiappeavis is a genus of enantiornithean bird from Early Cretaceous of northeastern China. The only species is Chiappeavis magnapremaxillo. Chiappeavis is classified within the family Pengornithidae. It is known from a single, almost complete skeleton including feather impressions discovered in the Jiufotang Formation of the Jehol Group. Long feathers formed a fan-shaped tail that was probably employed in flight.
Falcatakely is an extinct genus of enantiornithean bird known from partial fossils from northern Madagascar. The genus contains a single species, Falcatakely forsterae.
Yuanchuavis is an extinct genus of pengornithid bird in the group Enantiornithes. Its type and only specimen, IVPP V27883, is the holotype for its only species, Y. kompsosoura. It was recovered from the Jiufotang Formation of China, and has a fossilized fan of highly graduated rectrices. The tail has graduation similar to Chiappeavis and a pair of highly elongate central rectrices similar to the pintails of extant male sunbirds. Melanosomes sampled from the holotype suggest the central pair of rectrices to be dark in colour, and the rest of the tail feathers to be either grey or with non-iridescent structural colour.
