Cathayornis

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Cathayornis
Temporal range: Early Cretaceous, 120  Ma
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Cathayornis-Beijing Museum of Natural History.jpg
Specimen on display at the Beijing Museum of Natural History.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Avialae
Clade: Enantiornithes
Clade: Euenantiornithes
Genus: Cathayornis
Zhou, Jin & Zhang, 1992
Type species
Cathayornis yandica
Zhou et al., 1992
Other Species
  • C. aberransis? Hou et al., 2002
  • C. chabuensis? Li et al., 2008

Cathayornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known definitively from only one species, Cathayornis yandica, one of the first Enantiornithes found in China. Several additional species were once incorrectly classified as Cathayornis, and have since been reclassified or regarded as nomina dubia .

Contents

Description

Cathayornis yandica was a small enantiornithean with a slightly elongated, toothy snout and perching feet. Like most other Enantiornithes, it had large claws on the first two fingers that supported the wing. According to most recent studies, only one specimen can be definitively assigned to this species, a fossil catalogued as number IVPP V9769 and currently housed in the collections of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. [1] Cathayornis can be told apart from similar Enantiornithes (especially Sinornis , Eocathayornis , and Houornis ) by its larger size, a shorter and straighter first finger with a slightly longer claw, and other anatomical details. [2] Two additional but very fragmentary specimens, IVPP V9936 and V10896, have been referred to C. yandica in the past, but cannot be directly compared with the type specimen because they do not preserve any of the same key parts of the skeleton. [1]

Classification and species

Paul Sereno et al., in 2002, considered Cathayornis a junior synonym of Sinornis. They interpreted the anatomies of the two as very similar and sharing key autapomorphies of the pygostyle. [3] The first thorough review of Sinornis and Cathayornis was published by Jingmai O'Connor and Gareth Dyke in 2010. O'Connor and Dyke concluded that despite the earlier opinion of Sereno and colleagues, the two birds were not synonyms and in fact differ in several clear ways, including different proportions in the wing claws and digits, differences in the pelvis, and size of the pygostyle. [4]

Several other species Cathayornis aberransis, [5] Cathayornis chabuensis [6] and Cathayornis caudatus [7] had been classified as Cathayornis in the past. However, their validity and/or assignment to the genus Cathayornis has been questioned in subsequent evaluations. Jingmai O'Connor and Gareth Dyke (2010) found that many of the supposedly distinct features of C. aberransis (such as the base of a crest on the skull) had been inaccurately described, casting doubt on the few remaining features separating it from C. yandica, and suggested that further study was needed to determine its validity. [4] Similarly, C. caudatus was so named for its supposedly bony tail lacking a pygostyle, and was further differentiated by its small size. O'Connor and Dyke re-examined the specimen and showed that the specimen is in fact only slightly smaller than the type specimen of C. yandica, and that a normal enantiornithean tail with a pygostyle is clearly visible in one of the fossil slabs, parts of the hip bones having been mistaken for unfused tail vertebrae. O'Connor and Dyke therefore considered C. caudatus a nomen dubium. [4] They considered C. chabuenis, from the Jingchuan Formation of Inner Mongolia, to be clearly distinct from C. yandica and most likely a representative of a new genus. [4] In a 2015 re-evaluation of supposed "cathayornithids", Wang and Liu determined that C. caudatus could be differentiated from Cathayornis and placed it in the new genus Houornis . On the other hand, they considered C. chabuensis a nomen dubium. [1]

Other species of similar birds from the Jiufotang Formation have been regarded as synonymous with C. yandica by some researchers, including Largirostrornis sexdentoris and Cuspirostrisornis houi , [8] though this has yet to be supported by rigorous study. [4] O'Connor and colleagues noted that Longchengornis sanyanensis , also synonymized with C. yandica by some authors, seems to show distinct anatomy not shared with at least that species of Cathayornis. [4]

Related Research Articles

<i>Confuciusornis</i> Extinct genus of birds

Confuciusornis is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, Confuciusornis had a toothless beak, but closer and later relatives of modern birds such as Hesperornis and Ichthyornis were toothed, indicating that the loss of teeth occurred convergently in Confuciusornis and living birds. It was thought to be the oldest known bird to have a beak, though this title now belongs to an earlier relative Eoconfuciusornis. It was named after the Chinese moral philosopher Confucius. Confuciusornis is one of the most abundant vertebrates found in the Yixian Formation, and several hundred complete specimens have been found.

<span class="mw-page-title-main">Enantiornithes</span> Extinct clade of dinosaurs

The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.

<i>Yanornis</i> Extinct genus of birds

Yanornis is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.

<i>Longipteryx</i> Genus of birds

Longipteryx is a genus of prehistoric bird which lived during the Early Cretaceous. It contains a single species, Longipteryx chaoyangensis. Its remains have been recovered from the Jiufotang Formation at Chaoyang in Liaoning Province, China. Apart from the holotype IVPP V 12325 - a fine and nearly complete skeleton — another entire skeleton and some isolated bones are known to date.

<i>Sinornis</i> Extinct genus of birds

Sinornis is a genus of enantiornithean birds from the Lower Cretaceous Jiufotang Formation of the People's Republic of China.

<i>Cuspirostrisornis</i> Extinct genus of birds

Cuspirostrisornis is a genus of enantiornithean bird. Only one species is known, Cuspirostrisornis houi, though some researchers believe this to be a synonym of the similar species Cathayornis yandica. It is known from one fossil found in the Jiufotang Formation in Liaoning province, People's Republic of China. The Jiufotang Formation is dated to the Early Cretaceous period, Aptian age, 120.3 +/-0.7 million years ago.

Largirostrornis is a genus of enantiornithean bird. One species has been named, Largirostornis sexdentoris. It lived during the Early Cretaceous and is known from fossils found in the Jiufotang Formation in Liaoning province, People's Republic of China. Some researchers believe this species to be a junior synonym of the similar Cathayornis yandica.

<i>Liaoxiornis</i> Extinct genus of birds

Liaoxiornis is a dubious genus of enantiornithine bird. The only named species is Liaoxiornis delicatus, described by Hou and Chen in 1999. Because the species was named for a hatchling specimen, it cannot be matched with adult specimens, and so it is impossible to determine which, if any, birds from the same rocks represent adults of this species. Luis Chiappe and colleagues therefore regarded it as a nomen vanum or at least a nomen dubium, and recommended that use of the name be abandoned.

Longchengornis is a genus of enantiornithean birds which lived during the early Cretaceous Period, about 120 mya, and is known from a fossil found in the Jiufotang Formation in Liaoning province, People's Republic of China. It contains the single species L. sanyanensis, though some researchers believe this to be a synonym of the similar species Cathayornis yandica.

Neuquenornis volans is a species of enantiornithean birds which lived during the late Cretaceous period in today's Patagonia, Argentina. It is the only known species of the genus Neuquenornis. Its fossils were found in the Santonian Bajo de la Carpa Formation, dating from about 85-83 million years ago. This was a sizeable bird for its time, with a tarsometatarsus 46.8mm long. Informal estimates suggest that it measured nearly 30 cm (12 in) in length excluding the tail.

<i>Liaoningornis</i> Extinct genus of birds

Liaoningornis is a genus of bird from Lower Cretaceous China. It was collected from the dinosaur-bearing beds of the Sihetun locality, of the Yixian Formation, Shangyuan, near the city of Beipiao in Liaoning province. The only known species is Liaoningornis longidigitris. It was described by Linhai Hou in 1996 and 1997.

Paraprotopteryx is a genus of enantiornithean birds from the Mesozoic of China.

<i>Shanweiniao</i> Extinct genus of birds

Shanweiniao is a genus of long-snouted enantiornithean birds from Early Cretaceous China. One species is known, Shanweiniao cooperorum. There is one known fossil, a slab and counterslab. The fossil is in the collection of the Dalian Natural History Museum, and has accession number DNHM D1878/1 and DNHM1878/2. It was collected from the Lower Cretaceous Dawangzhengzi Beds, middle Yixian Formation, from Lingyuan in the Liaoning Province, China.

<span class="mw-page-title-main">Longipterygidae</span> Extinct family of dinosaurs

Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.

<i>Bohaiornis</i> Extinct genus of birds

Bohaiornis is a genus of enantiornithean birds. Fossils have been found from the Lower Cretaceous Jiufotang Formation of western Liaoning, China. The only known species, Bohaiornis guoi, was named by Dongyu Hu, Li Li, Lianhaim Hou and Xing Xu in 2011 on the basis of a fully articulated and well-preserved skeleton of a sub-adult. This specimen, LPM B00167, preserved two long, ribbon-like feathers attached to the tail rather than a fan of shorter pennaceous feathers. It was similar to the slightly older Eoenantiornis, but much larger in size. Bohaiornis is the type species of Bohaiornithidae, a family of large predatory enantiornitheans from the Early Cretaceous.

<span class="mw-page-title-main">Euornithes</span> Clade of dinosaurs

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Parabohaiornis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Parabohaiornis martini.

<i>Longusunguis</i> Extinct genus of birds

Longusunguis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Longusunguis kurochkini.

Houornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known from a single species, Houornis caudatus, which had been once been classified as a species of Cathayornis, and has also been regarded as a nomen dubium.

<i>Parapengornis</i> Extinct genus of birds

Parapengornis is an extinct genus of enantiornithine bird from the Lower Cretaceous of what is now China. The holotype specimen was discovered in the Jiufotang Formation near Lingyuan, western Liaoning province, and was catalogued as IVPP V18687. The nearly complete, articulated specimen is preserved on a slab and has impressions of pennaceous feathers. Only parts of the sternum, the left hand, and right foot are missing. In 2015, it became the basis of the new genus and species Parapengornis eurycaudatus, named by the Chinese palaeontologists Han Hu, Jingmai K. O’Connor, and Zhonghe Zhou. The generic name consists of the Latin word para and the name of the related genus Pengornis, indicating their close relationship. The name Pengornis is itself derived from "Peng", a mythological bird from Chinese folklore, and ornis, which means bird in Greek. The specific name is derived from the Latin words eury, meaning broad, and caudatus, meaning tail, in reference to the broad and expanded pygostyle. A nearly complete specimen formerly assigned to Pengornis was also reassigned to Parapengornis by these authors.

References

  1. 1 2 3 Wang, M.; Liu, D. (2015). "Taxonomical reappraisal of Cathayornithidae (Aves: Enantiornithes)". Journal of Systematic Palaeontology. 14: 29–47. doi:10.1080/14772019.2014.994087. S2CID   86665059.
  2. Zhou, Zhonghe, Hou and Lianhai. (2001). "The Discovery and Study of Mesozoic Birds in China." In Chiappe, L. and Witmer, L. (eds.), Mesozoic Birds: Above the Heads of Dinosaurs. 2001: University of California Press.
  3. Sereno, Rao and Li, (2002). "Sinornis santensis (Aves: Enantiornithes) from the Early Cretaceous of Northeastern China." Pp 184-208. in Chiappe and Witmer (eds.), Mesozoic Birds – Above the Heads of Dinosaurs. Berkeley: University of California Press
  4. 1 2 3 4 5 6 O'Connor, J. and Dyke, G. (2010). "A reassessment of Sinornis santensis and Cathayornis yandica (Aves: Enantiornithes)." Records of the Australian Museum, 62: 7-20. doi : 10.3853/J.0067-1975.62.2010.1540
  5. Hou, Zhou, Zhang and Gu, (2002). Mesozoic birds from western Liaoning in China. ISBN   7-5381-3392-5. 120 pp.
  6. Li, J., Li, Z., Zhang, Y., Zhou, Z., Bai, Z., Zhang, L. and Ba, T. (2008). "A new species of Cathayornis from the Lower Cretaceous of Inner Mongolia, China and its stratigraphic significance." Acta Geologica Sinica, 82(6): 1115-1123.
  7. Hou Lianhai, 1997. Mesozoic Birds of China. Phoenix Valley Bird Park, Lugu Hsiang, Taiwan. 221 pp.
  8. Zhou Z. and Wang Y. (2010). "Vertebrate diversity of the Jehol Biota as compared with other lagerstätten." Science China: Earth Sciences, 53(12): 1894–1907. doi : 10.1007/s11430-010-4094-9 Archived 2013-10-29 at the Wayback Machine