Houornis Temporal range: Early Cretaceous, | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Clade: | † Enantiornithes |
Genus: | † Houornis Wang & Liu 2015 |
Species: | †H. caudatus |
Binomial name | |
†Houornis caudatus (Hou, 1997) Wang & Liu 2015 | |
Synonyms | |
Genus synonymy
Species synonymy
|
Houornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known from a single species, Houornis caudatus, which had been once been classified as a species of Cathayornis , and has also been regarded as a nomen dubium .
Houornis caudatus was a small enantiornithean with a slightly elongated, toothy snout and perching feet. It is known only from a single specimen, the slab and counter-slab of a fossil catalogued as number IVPP V10917/2 in the collections of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. [1] The specimen was found in 1993 by Zhou Zhonghe near the town of Boluochi in Liaoning Province, China. The actual bones of this specimen were not preserved, but rather can be seen by the impressions they left in the surrounding rock. It was originally thought to have a relatively long, bony tail intermediate in length between modern birds and long-tailed birds like Archaeopteryx , and so was in 1997 by Hou Lianhu given the name Cathayornis caudatus, meaning "tailed Cathay bird". This interpretation was later found to be in error, though it does appear to have an unusually long pygostyle (a component of the tail made of fused vertebrae). [1]
While some researchers have considered the species dubious due to the poor preservation quality and incomplete nature of the fossil, a 2015 study by Wang Min and Liu Di was complete enough to be compared to similar species, including Cathayornis, and so they assigned it to its own genus, Houornis, the generic name honouring Hou. Wang and Liu found that Houornis can be told apart from similar species by several anatomical details, including the large pygostyle that gave it its name. [2] Specimen IVPP V10533, the rear of a skeleton, had in 1997 been referred to the species by Hou and this was confirmed in 2015. Two additional but very fragmentary specimens, IVPP V9936 and V10896, have been referred to C. yandica in the past, but cannot be directly compared with the type specimen because they do not preserve any of the same key parts of the skeleton. [1]
Cathayornis caudatus was so named for its supposedly long bony tail lacking a pygostyle, and was further differentiated by its small size (the humerus, or upper arm bone, is less than 30 millimetres (1.2 in) long). [1] In a 2010 paper, O'Connor and Dyke re-examined the specimen and showed that it is in fact only slightly smaller than the type specimen of C. yandica, and that a normal enantiornithean tail with a pygostyle is clearly visible in one of the fossil slabs, parts of the hip bones having been mistaken for unfused tail vertebrae. O'Connor and Dyke therefore considered C. caudatus a nomen dubium. [3] However, in a 2015 re-evaluation of supposed "cathayornithids", Wang and Liu determined that C. caudatus could be differentiated from Cathayornis, but could not determine its relationships with other members of Enantiornithes using a phylogenetic analysis. They placed Houornis as incertae sedis . [1]
Confuciusornis is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, Confuciusornis had a toothless beak, but closer and later relatives of modern birds such as Hesperornis and Ichthyornis were toothed, indicating that the loss of teeth occurred convergently in Confuciusornis and living birds. It was thought to be the oldest known bird to have a beak, though this title now belongs to an earlier relative Eoconfuciusornis. It was named after the Chinese moral philosopher Confucius. Confuciusornis is one of the most abundant vertebrates found in the Yixian Formation, and several hundred complete specimens have been found.
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Yanornis is an extinct genus of fish-eating Early Cretaceous birds. Two species have been described, both from Liaoning province, China: Yanornis martini, based on several fossils found in the 120-million-year-old Jiufotang Formation at Chaoyang, and Yanornis guozhangi, from the 124-million-year-old Yixian Formation.
Longipteryx is a genus of prehistoric bird which lived during the Early Cretaceous. It contains a single species, Longipteryx chaoyangensis. Its remains have been recovered from the Jiufotang Formation at Chaoyang in Liaoning Province, China. Apart from the holotype IVPP V 12325 - a fine and nearly complete skeleton — another entire skeleton and some isolated bones are known to date.
Cathayornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known definitively from only one species, Cathayornis yandica, one of the first Enantiornithes found in China. Several additional species were once incorrectly classified as Cathayornis, and have since been reclassified or regarded as nomina dubia.
Sinornis is a genus of enantiornithean birds from the Lower Cretaceous Jiufotang Formation of the People's Republic of China.
Cuspirostrisornis is a genus of enantiornithean bird. Only one species is known, Cuspirostrisornis houi, though some researchers believe this to be a synonym of the similar species Cathayornis yandica. It is known from one fossil found in the Jiufotang Formation in Liaoning province, People's Republic of China. The Jiufotang Formation is dated to the Early Cretaceous period, Aptian age, 120.3 +/-0.7 million years ago.
Largirostrornis is a genus of enantiornithean bird. One species has been named, Largirostornis sexdentoris. It lived during the Early Cretaceous and is known from fossils found in the Jiufotang Formation in Liaoning province, People's Republic of China. Some researchers believe this species to be a junior synonym of the similar Cathayornis yandica.
Liaoxiornis is a dubious genus of enantiornithine bird. The only named species is Liaoxiornis delicatus, described by Hou and Chen in 1999. Because the species was named for a hatchling specimen, it cannot be matched with adult specimens, and so it is impossible to determine which, if any, birds from the same rocks represent adults of this species. Luis Chiappe and colleagues therefore regarded it as a nomen vanum or at least a nomen dubium, and recommended that use of the name be abandoned.
Zhongornis is a genus of primitive avialan that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008.
Shanweiniao is a genus of long-snouted enantiornithean birds from Early Cretaceous China. One species is known, Shanweiniao cooperorum. There is one known fossil, a slab and counterslab. The fossil is in the collection of the Dalian Natural History Museum, and has accession number DNHM D1878/1 and DNHM1878/2. It was collected from the Lower Cretaceous Dawangzhengzi Beds, middle Yixian Formation, from Lingyuan in the Liaoning Province, China.
Dalingheornis is a genus of enantiornithean birds which lived during the early Cretaceous period, about 122 Ma ago, and are known from a single juvenile fossil found in the upper part of the Yixian Formation at Dawangzhangzi, Liaoning province, People's Republic of China. It is the first known Mesozoic bird with heterodactyl feet specifically adapted for climbing, and was probably among the most arboreal of the enantiornithines. Unlike its relatives, it had an unusually long (17mm) skeletal tail made up of 20 vertebrae, similar to the tails of dromaeosaurids. However, this may have been a juvenile feature. The fossil was named after Yang Liwei, the first Chinese astronaut in space.
Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.
Bohaiornis is a genus of enantiornithean birds. Fossils have been found from the Lower Cretaceous Jiufotang Formation of western Liaoning, China. The only known species, Bohaiornis guoi, was named by Dongyu Hu, Li Li, Lianhaim Hou and Xing Xu in 2011 on the basis of a fully articulated and well-preserved skeleton of a sub-adult. This specimen, LPM B00167, preserved two long, ribbon-like feathers attached to the tail rather than a fan of shorter pennaceous feathers. It was similar to the slightly older Eoenantiornis, but much larger in size. Bohaiornis is the type species of Bohaiornithidae, a family of large predatory enantiornitheans from the Early Cretaceous.
Parabohaiornis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Parabohaiornis martini.
Longusunguis is an extinct genus of bohaiornithid enantiornithean dinosaur known from the Early Cretaceous of Liaoning Province, northeastern China. It contains a single species, Longusunguis kurochkini.
Cruralispennia is an extinct genus of enantiornithean bird. The only known specimen of Cruralispennia was discovered in the Early Cretaceous Huajiying Formation of China and formally described in 2017. The type species of Cruralispennia is Cruralispennia multidonta. The generic name is Latin for "shin feather", while the specific name means "many-toothed". The holotype of Cruralispennia is IVPP 21711, a semi-articulated partial skeleton surrounded by the remains of carbonized feathers.
Parapengornis is an extinct genus of enantiornithine bird from the Lower Cretaceous of what is now China. The holotype specimen was discovered in the Jiufotang Formation near Lingyuan, western Liaoning province, and was catalogued as IVPP V18687. The nearly complete, articulated specimen is preserved on a slab and has impressions of pennaceous feathers. Only parts of the sternum, the left hand, and right foot are missing. In 2015, it became the basis of the new genus and species Parapengornis eurycaudatus, named by the Chinese palaeontologists Han Hu, Jingmai K. O’Connor, and Zhonghe Zhou. The generic name consists of the Latin word para and the name of the related genus Pengornis, indicating their close relationship. The name Pengornis is itself derived from "Peng", a mythological bird from Chinese folklore, and ornis, which means bird in Greek. The specific name is derived from the Latin words eury, meaning broad, and caudatus, meaning tail, in reference to the broad and expanded pygostyle. A nearly complete specimen formerly assigned to Pengornis was also reassigned to Parapengornis by these authors.
Chiappeavis is a genus of enantiornithean bird from Early Cretaceous of northeastern China. The only species is Chiappeavis magnapremaxillo. Chiappeavis is classified within the family Pengornithidae. It is known from a single, almost complete skeleton including feather impressions discovered in the Jiufotang Formation of the Jehol Group. Long feathers formed a fan-shaped tail that was probably employed in flight.
Avimaia is a genus of fossil birds of the Enantiornithes clade that lived about 115 million years ago in Northwest China. The only known species is A. schweitzerae. The holotype fossil of the bird was found in the Xiagou Formation, and is noted as the first discovered fossil bird with an unlaid egg. Abnormalities, including egg binding in which the egg becomes stuck within the body of the bird causing death, were found in the egg suggesting that the preserved egg may have caused this bird's demise. Egg binding is a serious and lethal condition that is fairly common in small birds undergoing stress.