Feitianius Temporal range: Early Cretaceous, | |
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Life restoration of Feitianius paradisi. | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Clade: | Pygostylia |
Clade: | Ornithothoraces |
Clade: | † Enantiornithes |
Genus: | † Feitianius O'Connor et al., 2015 |
Type species | |
Feitianius paradisi O'Connor et al., 2015 |
Feitianius is a bird genus, belonging to the Enantiornithes, that during the Early Cretaceous lived in the area of modern China. A single species has been named in the genus, Feitianius paradisi.
In 2005, fragmentary enantiornithean fossils were excavated near Changma, in Gansu. Two of the best preserved were reported in the scientific literature in 2009. [1]
In 2015, the type species Feitianius paradisi was named and described by Jingmai Kathleen O’Connor, Li Daqing, Matthew C. Lamanna, Wang Min, Jerald D. Harris, Jessie Atterholt and You Hailou. The generic name has been derived from the feitia, a flying apsara or nymph from Buddhist mythology. Historical murals featuring such nymphs are common in the caves around Changma. The name was "for fun" made masculine because it was concluded that the type specimen was a male individual. The specific name means "from paradise" in Latin and refers to the similar sexual dimorphism in the tail feathers of birds-of-paradise. The species name as a whole thus means "feitia of paradise", the translation of the Chinese 飛天/飞天. [2]
The holotype, GSGM-05-CM-004, was found in a layer of the Xiagou Formation dating from the early Aptian. In consists of the hind part of a skeleton on a stone plate. The skeleton is visible from the left side. It is articulated but the part in front of the middle back is lacking. It preserves extensive remains of the soft tissues, including the plumage. [2]
An indication of the size of Feitianius is given by the length of the thighbone, twenty-seven millimetres. [2]
A single unique feature was established for Feitianius: the tail plumage contains three distinct feather types whereas all known relatives have tails with a single type of feather. [2]
Soft tissue remains are preserved around the tail base. These seem to represent a fleshy structure similar to the "parson's nose" of modern birds, in which the tail feathers are embedded. Collagen bands are visible running perpendicular to the body outline. These bands have a length of seven to eight millimetres and a width of about half a millimetre. [2]
On the belly and the front tail base short downy covert feathers are visible. More to behind the feathers of the tail base are gradually changing into longer structures, a dozen of which are clearly visible. Uniquely for the Enantiornithes, among these three morphotypes can be distinguished. Morphotype A consists of feathers that are forty to forty-five millimetres long. They have vanes that cover the distal 55% of the rachis. Morphotype B is eighteen millimetres long and half as wide as morphotype A. Here, the vanes cover the full length of the rachis. Both types are pennaceous and somewhat curved downwards. Of morphotype C only a single rachis has been preserved. It is very long and its distal end has been lost on the fossil. The describing authors concluded that it was one of a pair of very long central "streamers" having a display function, probably for a male individual. It was considered unlikely that the tail feathers had an important aerodynamic effect. It is possible that the shorter morphotype A and B feathers formed a differentially coloured tuft on the tail base. Because of a varnish applied to the fossil, it can no longer be checked for colouring melanosomes though. The ornamental structure as a whole might have resembled that of the extant greater bird-of-paradise. [2]
Feitianius was in 2016 placed in the Enantiornithes. A cladistic analysis showed it to be part of a polytomy with Qiliania and other enantiornithean specimens of the Changma Basin. [2]
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Iberomesornis is a monotypic genus of enantiornithine bird of the Cretaceous of Spain.
Gansus is a genus of aquatic birds that lived during the Aptian age of the Early Cretaceous (Aptian-Albian) period in what are now Gansu and Liaoning provinces, western China. The rock layers from which their fossils have been recovered are dated to 120 million years ago. It was first described in 1984 on the basis of an isolated left leg. It is the oldest-known member of the Ornithurae, the group which includes modern birds (Neornithes) and extinct related groups, such as Ichthyornis and Hesperornithes.
The Xiagou Formation is the middle strata of the Xinminbao Group. It is named for its type site in Xiagou, in the Changma Basin of Gansu Province, northwestern China and is considered Early Cretaceous in age. It is known outside the specialized world of Chinese geology as the site of a Lagerstätte in which the fossils were preserved of Gansus yumenensis, the earliest true modern bird.
Eoalulavis is a monotypic genus of enantiornithean bird that lived during the Barremian, in the Lower Cretaceous around 125 million years ago. The only known species is Eoalulavis hoyasi.
Paraprotopteryx is a genus of enantiornithean birds from the Mesozoic of China.
Protopteryx is an extinct bird and the most basal enantiornithean, from the Cretaceous period. The type species is P. fengningensis. It was first discovered in the Sichakou Member of the Yixian Formation or Huajiying Formation of Hebei Province, northern China, dating from 131 Ma ago. Protopteryx has been found in the Daibeigou formation, as well. The name Protopteryx means "primitive feather": "proto-" meaning "the first of" and "-pteryx" meaning "feather" or "wing." The name comes from the fact that Protopteryx feathers are more primitive than those of modern birds, such as the two elongated tail feathers that lack barbs and rami.
Avisauridae is a family of extinct enantiornithine dinosaurs from the Cretaceous period, distinguished by several features of their ankle bones. Depending on the definition used, Avisauridae is either a broad and widespread group of advanced enantiornithines, or a small family within that group, restricted to species from the Late Cretaceous of North and South America.
Shanweiniao is a genus of long-snouted enantiornithean birds from Early Cretaceous China. One species is known, Shanweiniao cooperorum. There is one known fossil, a slab and counterslab. The fossil is in the collection of the Dalian Natural History Museum, and has accession number DNHM D1878/1 and DNHM1878/2. It was collected from the Lower Cretaceous Dawangzhengzi Beds, middle Yixian Formation, from Lingyuan in the Liaoning Province, China.
Bohaiornis is a genus of enantiornithean birds. Fossils have been found from the Lower Cretaceous Jiufotang Formation of western Liaoning, China. The only known species, Bohaiornis guoi, was named by Dongyu Hu, Li Li, Lianhaim Hou and Xing Xu in 2011 on the basis of a fully articulated and well-preserved skeleton of a sub-adult. This specimen, LPM B00167, preserved two long, ribbon-like feathers attached to the tail rather than a fan of shorter pennaceous feathers. It was similar to the slightly older Eoenantiornis, but much larger in size. Bohaiornis is the type species of Bohaiornithidae, a family of large predatory enantiornitheans from the Early Cretaceous.
Qiliania is an extinct genus of early bird from the lower Cretaceous about 120 million years ago. It is an enantiornithean which lived in what is now Gansu Province, north-western China. It is known from two incomplete, semi-articulated and three-dimensionally preserved skeletons, which were found in the Xiagou Formation of the Changma Basin. It was first named by Shu-An Ji, Jessie Atterholt, Jingmai O'Connor, Matthew Lamanna, Jerry Harrs, Li Da-Qing, You Hai-Lu and Peter Dodson in 2011 and the type species is Qiliania graffini. The species was named for Greg Graffin, a paleontologist and member of the punk rock group Bad Religion, at the suggestion of coauthor Jingmai O'Connor, a longtime fan of the band.
Houornis is a genus of enantiornithean birds from the Jiufotang Formation of Liaoning, People's Republic of China. It is known from a single species, Houornis caudatus, which had been once been classified as a species of Cathayornis, and has also been regarded as a nomen dubium.
Pengornithidae is a group of early enantiornithines from the early Cretaceous Period of China, with the putative member Falcatakely possibly extending this clade's range into the Late Cretaceous of Madagascar, and several putative pengornithids also hail from this formation. Specimens of these animals have been found both in the Huajiying Formation and Jiufotang Formation of Liaoning and Hebei provinces, dating from the Hauterivian age to the Aptian age.
Dunhuangia is a bird genus, belonging to the Enantiornithes, which during the Early Cretaceous lived in the area of present China.
Cruralispennia is an extinct genus of enantiornithean bird. The only known specimen of Cruralispennia was discovered in the Early Cretaceous Huajiying Formation of China and formally described in 2017. The type species of Cruralispennia is Cruralispennia multidonta. The generic name is Latin for "shin feather", while the specific name means "many-toothed". The holotype of Cruralispennia is IVPP 21711, a semi-articulated partial skeleton surrounded by the remains of carbonized feathers.
Avimaia is a genus of fossil birds of the Enantiornithes clade that lived about 115 million years ago in Northwest China. The only known species is A. schweitzerae. The holotype fossil of the bird was found in the Xiagou Formation, and is noted as the first discovered fossil bird with an unlaid egg. Abnormalities, including egg binding in which the egg becomes stuck within the body of the bird causing death, were found in the egg suggesting that the preserved egg may have caused this bird's demise. Egg binding is a serious and lethal condition that is fairly common in small birds undergoing stress.
Grabauornis is an extinct genus of enantiornitheans from the Early Cretaceous of China.
Jingmai Kathleen O'Connor is a paleontologist who works as a curator at the Field Museum.
Yuanjiawaornis is an extinct genus of large enantiornithean bird known from the early Cretaceous of present-day China. It is monotypic, with only type species Y. virisosus known.