Caenagnathasia Temporal range: Turonian-Maastrichtian, | |
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Illustration of the lower jaw | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Caenagnathidae |
Subfamily: | † Elmisaurinae |
Genus: | † Caenagnathasia Currie et al., 1994 |
Species: | †C. martinsoni |
Binomial name | |
†Caenagnathasia martinsoni Currie et al., 1994 | |
Caenagnathasia ('recent jaw from Asia') is a small caenagnathid oviraptorosaurian theropod from the Late Cretaceous of Uzbekistan.
The type species Caenagnathasia martinsoni was named and described in 1994 by Philip J. Currie, Stephen Godfrey and Lev Nesov. The generic name is a combination of a reference to the species' placement in the Caenagnathidae, with Asia, the continent of its provenance. The specific name honours Gerbert Genrikhovich Martinson. The genus is based on holotype N 401/12457, a pair of fused dentaries of the lower jaws. The specimen was found near Dzharakuduk in layers of the Bissekty Formation, dating to the Turonian-Coniacian, around 90 mya, making it the oldest known caenagnathoid. A second specimen was referred to the species, N 402/12457, a right dentary of a slightly smaller individual. Both individuals were adult. [1] In 2015, new material of Caenagnathasia was described. From the same site as the holotype, the material includes various vertebrae, a dentary, and a femur. [2]
Caenagnathasia is the smallest known oviraptorosaur and one of the smallest non-avian dinosaurs. The jaw fragments are only a few centimetres long and total skull length has been estimated at 3 inches (7.6 cm). A 2010 estimate by Gregory S. Paul gave it a length of 2 feet (0.61 m) and a weight of 3 pounds (1.4 kg). [3] Caenagnathasia would presumably have resembled other oviraptorosaurs, which were feathered, bird-like dinosaurs with beaked skulls, long necks, and long limbs. Recent studies suggest it was one of the more primitive members of the Caenagnathidae.
Caenagnathasia was originally assigned to the Caenagnathidae. It was even included in the definition of that clade published by Hans-Dieter Sues. [4] It has been suggested however, that it might have a more basal position in the Oviraptorosauria, outside of the Caenagnathoidea. [5] In 2015 after the description of new material, it was found that Caenagnathasia could confidently be referred to Caenagnathidae. [2]
The below cladogram is based on that resolved in the description of Anzu . [6]
Caenagnathoidea |
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Oviraptorosaurs are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from Caudipteryx, which was the size of a turkey, to the 8-meter-long, 1.4-ton Gigantoraptor. The group is close to the ancestry of birds. Some researchers such as Maryanska et al (2002) and Osmólska et al. (2004) have proposed that they may represent primitive flightless birds. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.
Nomingia is a genus of oviraptorid theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia.
Chirostenotes is a genus of oviraptorosaurian dinosaur from the late Cretaceous of Alberta, Canada. The type species is Chirostenotes pergracilis.
Conchoraptor is a genus of oviraptorid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 70 million years ago. It is known from the Barun Goyot and Nemegt formations of Mongolia.
Hagryphus is a monospecific genus of caenagnathid dinosaur from southern Utah that lived during the Late Cretaceous in what is now the Kaiparowits Formation of the Grand Staircase–Escalante National Monument. The type and only species, Hagryphus giganteus, is known only from an incomplete but articulated left manus and the distal portion of the left radius. It was named in 2005 by Lindsay E. Zanno and Scott D. Sampson. Hagryphus has an estimated length of 2.4–3 metres and weight of 50 kilograms.
Caenagnathus is a genus of caenagnathid oviraptorosaurian dinosaur from the late Cretaceous period. It is known from partial remains including lower jaws, a tail vertebra, hand bones, and hind limbs, all found in the Dinosaur Park Formation of Alberta, Canada. Caenagnathus measured about 2.5 m (8.2 ft) long and weighed about 96–100 kg (212–220 lb).
Elmisaurus is an extinct genus of caenagnathid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was a theropod belonging to the Oviraptorosauria.
Caenagnathidae is a family of derived caenagnathoid dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that Chirostenotes pergracilis, originally named on the basis of a pair of hands, and Citipes elegans, originally thought to be an ornithomimid, named from a foot, were caenagnathids as well.
Urbacodon is a genus of troodontid dinosaur, a type of small carnivore. It lived in Uzbekistan during the early Late Cretaceous Period, about 95 million years ago.
Gigantoraptor is a genus of large oviraptorosaur dinosaur that lived in Asia during the Late Cretaceous period. It is known from the Iren Dabasu Formation of Inner Mongolia, where the first remains were found in 2005.
The Iren Dabasu Formation is a Late Cretaceous geologic formation in the Iren Nor region of Inner Mongolia. Dinosaur remains diagnostic to the genus level are among the fossils that have been recovered from the formation. The formation was first described and defined by Henry Fairfield Osborn in 1922 and it is located in the Iren Nor region of China.
Nankangia is an extinct genus of caenagnathoid oviraptorosaurian dinosaur known from the Upper Cretaceous Nanxiong Formation of Nankang County, Ganzhou City of Jiangxi Province, southeastern China. It contains a single species, Nankangia jiangxiensis. N. jiangxiensis coexisted with at least four other caenagnathoids, including but not limited to Corythoraptor, Banji, Ganzhousaurus and Jiangxisaurus. The relatively short dentary and non-downturned mandibular symphysis of Nankangia suggest that it may have been more herbivorous than carnivorous. Its diet consisted of leaves and seeds.
Anzu is a monospecific genus of caenagnathid dinosaur from North Dakota, South Dakota and Montana that lived during the Late Cretaceous in what is now the Hell Creek Formation. The type species and only species, Anzu wyliei is known from numerous skeletons that preserve cranial and postcranial elements. It was named in 2014 by Matthew C. Lamanna, Hans-Dieter Sues, Emma R. Schachner, and Tyler R. Lyson.
This timeline of oviraptorosaur research is a chronological listing of events in the history of paleontology focused on the oviraptorosaurs, a group of beaked, bird-like theropod dinosaurs. The early history of oviraptorosaur paleontology is characterized by taxonomic confusion due to the unusual characteristics of these dinosaurs. When initially described in 1924 Oviraptor itself was thought to be a member of the Ornithomimidae, popularly known as the "ostrich" dinosaurs, because both taxa share toothless beaks. Early caenagnathid oviraptorosaur discoveries like Caenagnathus itself were also incorrectly classified at the time, having been misidentified as birds.
Timurlengia is an extinct genus of tyrannosauroid theropod dinosaur found in Uzbekistan, in the Bissekty Formation in the Kyzylkum Desert, hailing from the Turonian age of the early Late Cretaceous. The type species is Timurlengia euotica.
Apatoraptor is a genus of caenagnathid dinosaur which contains a single species, A. pennatus. The only known specimen was discovered in the Campanian-age Horseshoe Canyon Formation of Alberta.
Beibeilong is a genus of large caenagnathid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 96 million to 88 million years ago. The genus contains a single species, Beibeilong sinensis. The species was named and described in 2017 through analysis of an embryonic skeleton and partial nest with large eggs that were discovered in the Gaogou Formation of China between 1992 and 1993.
Xingtianosaurus is an extinct genus of oviraptorosaurian theropod dinosaur that lived in what is now China during the Early Cretaceous. The type and only species, X. ganqi, was named and described in 2019. It was placed in the Caudipteridae, alongside Caudipteryx and Similicaudipteryx.
Citipes is an extinct genus of caenagnathid theropod from the Late Cretaceous Dinosaur Park Formation in Alberta, Canada. The genus contains only one species, the type species, C. elegans. The generic name of Citipes is Latin for "fleet-footed", and the specific epithet "elegans" is Latin for "elegant". The type specimen of Citipes has a convoluted taxonomic history, and has been previously assigned to the genera Ornithomimus, Macrophalangia, Elmisaurus, Chirostenotes, and Leptorhynchos before being given its own genus in 2020.