Conchoraptor Temporal range: Late Cretaceous, ~ | |
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Cast mounted in a nesting position | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Oviraptoridae |
Subfamily: | † Heyuanninae |
Genus: | † Conchoraptor Barsbold, 1986 |
Species: | †C. gracilis |
Binomial name | |
†Conchoraptor gracilis Barsbold, 1986 | |
Conchoraptor (meaning "conch plunderer") is a genus of oviraptorid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 70 million years ago. It is known from the Barun Goyot and Nemegt formations of Mongolia.
When first discovered in the Red Beds of Hermiin Tsav of the Early Maastrichtian Nemegt Formation by a Polish-Mongolian expedition in 1971, scientists believed that Conchoraptor was a juvenile Oviraptor and that the animal's missing crest would have begun to grow when the animal reached sexual maturity. Further study of multiple skeletons showed that Conchoraptor belonged to a new genus. The hands of Conchoraptor were a major reason scientists decided to split it off from Oviraptor. Anatomically the hands seemed to be an evolutionary intermediate between those of Ajancingenia and Oviraptor, making it obvious that this animal was not a member of a known species. [1]
The type species of this new genus, Conchoraptor gracilis, was described and named by Barsbold, in 1986. The generic name is derived from Greek konkhè, "mussel", and Latin raptor, "plunderer". [2] This name reflects Barsbold's hypothesis that oviraptorids, rather than preying primarily upon eggs as had been traditionally thought, may have been specialized to feed on mollusks. [1] The specific name means "gracile" in Latin. The holotype is MPC-D 100/20, a partial skull. [3] About two dozen specimens were formally referred to the species later.
Conchoraptor was a small dinosaur, only 1–2 meters (3.3-6.6 feet) in length. Unlike many other oviraptorids, Conchoraptor lacked a head crest. Like its relatives, it lacked teeth. Instead of teeth, oviraptorids had powerful beaks, possibly adapted to crushing mollusk shells. [1] The skull of Conchoraptor was heavily pneumatised, with most of the bones hollow. [4]
Conchoraptor was by Barsbold assigned to the Oviraptoridae in 1986. Recent analyses show it was a member of the oviraptorid subfamily Ingeniinae (now Heyuanninae). The cladogram below follows an analysis by Fanti et al., 2012. [5]
Oviraptoridae |
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Internal connections between both ears functioned as resonance chambers and improved hearing. [6]
Originally it was suggested that Conchoraptor had a diet that primarily consisted of mollusks. [7] [8] [9] A 2022 study of the bite force of Conchoraptor and comparisons with other oviraptorosaurs such as Incisivosaurus , Citipati , and Khaan instead suggests that Conchoraptor had a very strong bite force. The moderate jaw gape seen in oviraptorosaurs is indicative of herbivory in the majority of the group, but it is clear they were likely feeding on much tougher or more various types of vegetation than other herbivorous theropods in their environment, such as ornithomimosaurs and therizinosaurs were able to. The examinations suggest oviraptorosaurs may have been powerful-biting generalists or specialists that partook of niche partitioning both in body size and cranial function. [10]
The Nemegt Formation of Mongolia, is estimated to date back to the Maastrichtian stage, between 70 and 66 million years ago [11] of the Late Cretaceous period. [12] During the Late Cretaceous period, the land that is now the Nemegt Formation had an arid environment with fields of sand dunes and only intermittent streams. It is slightly younger than the nearby Djadochta Formation, and seems to have been slightly wetter. [12] This formation is noteworthy for the exquisite preservation of small and fragile dinosaur skeletons, a rare occurrence considering that these fossils are typically broken up and dispersed when found in other rock formations. [13]
Conchoraptor shared its habitat with other oviraptorosaurs including Avimimus and Nomingia , other theropods include troodontids such as Tochisaurus and Zanabazar , the tyrannosaurids Alioramus and Tarbosaurus . Ornithischians include the pachycephalosaurs Homalocephale and Prenocephale , the hadrosaurid Saurolophus , the ankylosaurs Saichania and Tarchia .
Oviraptor is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period. The first remains were collected from the Djadokhta Formation of Mongolia in 1923 during a paleontological expedition led by Roy Chapman Andrews, and in the following year the genus and type species Oviraptor philoceratops were named by Henry Fairfield Osborn. The genus name refers to the initial thought of egg-stealing habits, and the specific name was intended to reinforce this view indicating a preference over ceratopsian eggs. Despite the fact that numerous specimens have been referred to the genus, Oviraptor is only known from a single partial skeleton regarded as the holotype, as well as a nest of about fifteen eggs and several small fragments from a juvenile.
Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one and two metres long in most cases, though some possible oviraptorids were enormous. Oviraptorids are currently known only from the Late Cretaceous of Asia, with the most well-known species and complete specimens found only in the Gobi Desert of Mongolia and northwestern China.
Khaan was an oviraptorid dinosaur that was found in the Djadochta Formation of Mongolia and lived in the Late Cretaceous Period (Campanian), 75-71 million years ago.
Avimimus, meaning "bird mimic", is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 to 70 million years ago.
Nomingia is a genus of oviraptorid theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia.
Citipati is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality at the Djadochta Formation, where the first remains were collected during the 1990s. The genus and type species Citipati osmolskae were named and described in 2001. A second species from the adjacent Zamyn Khondt locality may also exist. Citipati is one of the best-known oviraptorids thanks to a number of well-preserved specimens, including individuals found in brooding positions atop nests of eggs, though most of them were initially referred to the related Oviraptor. These nesting specimens have helped to solidify the link between non-avian dinosaurs and birds.
Heyuannia is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous epoch, in what is now China and Mongolia. It was the first oviraptorid found in China; most others were found in neighbouring Mongolia. Two species are known: H. huangi, named by Lü Junchang in 2002 from the Dalangshan Formation; and H. yanshini, originally named as a separate genus Ingenia from the Barun Goyot Formation by Rinchen Barsbold in 1981, and later renamed to Ajancingenia in 2013 due to the preoccupation of Ingenia. The latter name was eventually discarded due to various ethical issues surrounding the author.
Nemegtomaia is a genus of oviraptorid dinosaur from what is now Mongolia that lived in the Late Cretaceous Period, about 70 million years ago. The first specimen was found in 1996, and became the basis of the new genus and species N. barsboldi in 2004. The original genus name was Nemegtia, but this was changed to Nemegtomaia in 2005, as the former name was preoccupied. The first part of the generic name refers to the Nemegt Basin, where the animal was found, and the second part means "good mother", in reference to the fact that oviraptorids are known to have brooded their eggs. The specific name honours the palaeontologist Rinchen Barsbold. Two more specimens were found in 2007, one of which was found on top of a nest with eggs, but the dinosaur had received its genus name before it was found associated with eggs.
Rinchenia is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous epoch in what is now Mongolia, Nemegt Formation, around 70 million years ago. The type and only known species, Rinchenia mongoliensis, was originally classified as a species within the genus Oviraptor, but a subsequent reexamination found differences significant enough to warrant a separate genus. The name Rinchenia was coined for this new genus, though not formally described in detail.
The Barun Goyot Formation is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia.
The Nemegt Formation is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds.
Caenagnathidae is a family of derived caenagnathoid dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that Chirostenotes pergracilis, originally named on the basis of a pair of hands, and Citipes elegans, originally thought to be an ornithomimid, named from a foot, were caenagnathids as well.
Machairasaurus is a genus of oviraptorid dinosaur which was found in the Bayan Mandahu Formation, China, dating to the late Cretaceous period.
Macroolithus is an oogenus of dinosaur egg belonging to the oofamily Elongatoolithidae. The type oospecies, M. rugustus, was originally described under the now-defunct oogenus name Oolithes. Three other oospecies are known: M. yaotunensis, M. mutabilis, and M. lashuyuanensis. They are relatively large, elongated eggs with a two-layered eggshell. Their nests consist of large, concentric rings of paired eggs. There is evidence of blue-green pigmentation in its shell, which may have helped camouflage the nests.
Nankangia is an extinct genus of caenagnathoid oviraptorosaurian dinosaur known from the Upper Cretaceous Nanxiong Formation of Nankang County, Ganzhou City of Jiangxi Province, southeastern China. It contains a single species, Nankangia jiangxiensis. N. jiangxiensis coexisted with at least four other caenagnathoids, including but not limited to Corythoraptor, Banji, Ganzhousaurus and Jiangxisaurus. The relatively short dentary and non-downturned mandibular symphysis of Nankangia suggest that it may have been more herbivorous than carnivorous. Its diet consisted of leaves and seeds.
This timeline of oviraptorosaur research is a chronological listing of events in the history of paleontology focused on the oviraptorosaurs, a group of beaked, bird-like theropod dinosaurs. The early history of oviraptorosaur paleontology is characterized by taxonomic confusion due to the unusual characteristics of these dinosaurs. When initially described in 1924 Oviraptor itself was thought to be a member of the Ornithomimidae, popularly known as the "ostrich" dinosaurs, because both taxa share toothless beaks. Early caenagnathid oviraptorosaur discoveries like Caenagnathus itself were also incorrectly classified at the time, having been misidentified as birds.
Elongatoolithidae is an oofamily of fossil eggs, representing the eggs of oviraptorosaurs. They are known for their highly elongated shape. Elongatoolithids have been found in Europe, Asia, and both North and South America.
Ondogurvel is a genus of alvarezsaurid dinosaur from the Late Cretaceous (Campanian) Barun Goyot Formation in southern Mongolia. The type and only species is O. alifanovi, known from a partial skeleton consisting of fragments of two last dorsal vertebrae, three anterior sacral vertebrae, right ilium, left and right pubis and ischium, articulated right tibia, fibula, metatarsals II and IV, and phalanges IV-1 and IV-2, right carpometacarpus, left and right manual phalanx II-1, right femur, left pedal phalanx II-1, and fragments of unidentified phalanges.
Natovenator is a genus of halszkaraptorine dromaeosaurid dinosaur from the Late Cretaceous Barun Goyot Formation of Mongolia. The genus is known from a single species, N. polydontus. Natovenator is crucial to the understanding of halszkaraptorines due to it providing more support for the semi-aquatic lifestyle that has been proposed for this clade. This discovery is important as the semi-aquatic lifestyles of halszkaraptorines was contested in early 2022.
Jaculinykus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Baruungoyot Formation of Mongolia. The genus contains a single species, J. yaruui, known from a nearly complete articulated skeleton including bones of the skull. Jaculinykus is notable for its unique hand, which has a hypertrophied first digit and greatly reduced second digit, which is intermediate between the tridactyl hand of Shuvuuia and monodactyl hand of Linhenykus.