Caenagnathoidea

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Caenagnathoidea
Temporal range: Cretaceous, 125–66  Ma
Khaan.jpg
Fossil skeleton of Khaan mckennai
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Oviraptorosauria
Superfamily: Caenagnathoidea
Sternberg, 1940
Type species
Caenagnathus collinsi
Sternberg, 1940
Subgroups
Synonyms

Caenagnathoidea ("recent jaw forms") is a group of advanced oviraptorosaurian dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, often with bony crests atop the head. They ranged in size from Caudipteryx , which was the size of a turkey, to the 8 meter long, 1.4 ton Gigantoraptor . [1] The group (along with all maniraptoran dinosaurs) is close to the ancestry of birds. The most complete specimens have been found in Asia, representing members of the sub-group Oviraptorinae. [2] Notable but fragmentary remains are also known from North America, almost all of which belong to the subgroup Elmisaurinae. [2]

The earliest definitive caenagnathoid is Microvenator celer , which dates to the late Aptian age of the early Cretaceous period, though the slightly earlier Caudipteryx from the lower Yixian Formation of China, may also be a member of this group.

Classification

The internal classification of the oviraptorosaurs has also been controversial. Most early studies divided them into two primary sub-groups, the Caenagnathidae and the Oviraptoridae. The Oviraptoridae was further divided into the small, short-armed, and crestless subfamily Ingeniinae, and the larger, crested, long-armed Oviraptorinae. However, later phylogenetic studies showed that many traditional members of the Caenagnathidae were in fact more closely related to the crested oviraptorids. Because Caenagnathus itself is no longer considered part of this group, the subfamily is usually now termed the Elmisaurinae. The taxonomy here follows the one presented by Thomas Holtz, Jr. in 2010 unless otherwise noted. [3]

The following cladogram follows an analysis by Phil Senter, 2007. [4]

Caenagnathoidea

See also

Related Research Articles

<span class="mw-page-title-main">Oviraptoridae</span> Extinct family of dinosaurs

Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one and two metres long in most cases, though some possible oviraptorids were enormous. Oviraptorids are currently known only from the Late Cretaceous of Asia, with the most well-known species and complete specimens found only in the Gobi Desert of Mongolia and northwestern China.

<span class="mw-page-title-main">Alvarezsauridae</span> Family of extinct long-legged dinosaurs

Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests.

<span class="mw-page-title-main">Ornithomimosauria</span> Extinct clade of theropod dinosaurs

Ornithomimosauria are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia, as well as Africa and possibly Australia. The group first appeared in the Early Cretaceous and persisted until the Late Cretaceous. Primitive members of the group include Nqwebasaurus, Pelecanimimus, Shenzhousaurus, Hexing and Deinocheirus, the arms of which reached 2.4 m (8 feet) in length. More advanced species, members of the family Ornithomimidae, include Gallimimus, Struthiomimus, and Ornithomimus. Some paleontologists, like Paul Sereno, consider the enigmatic alvarezsaurids to be close relatives of the ornithomimosaurs and place them together in the superfamily Ornithomimoidea.

<span class="mw-page-title-main">Oviraptorosauria</span> Extinct clade of dinosaurs

Oviraptorosaurs are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from Caudipteryx, which was the size of a turkey, to the 8-meter-long, 1.4-ton Gigantoraptor. The group is close to the ancestry of birds. Some researchers such as Maryanska et al (2002) and Osmólska et al. (2004) have proposed that they may represent primitive flightless birds. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.

<i>Avimimus</i> Extinct genus of reptiles

Avimimus, meaning "bird mimic", is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 to 70 million years ago.

<i>Caenagnathasia</i> Extinct species of reptile

Caenagnathasia is a small caenagnathid oviraptorosaurian theropod from the Late Cretaceous of Uzbekistan.

<i>Nomingia</i> Extinct genus of dinosaurs

Nomingia is a genus of oviraptorosaur theropod dinosaur hailing from the Late Cretaceous Bugin Tsav Beds of Mongolia.

<i>Chirostenotes</i> Extinct genus of dinosaurs

Chirostenotes is a genus of oviraptorosaurian dinosaur from the late Cretaceous of Alberta, Canada. The type species is Chirostenotes pergracilis.

<i>Conchoraptor</i> Extinct genus of dinosaurs

Conchoraptor is a genus of oviraptorid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 70 million years ago. It is known from the Barun Goyot and Nemegt formations of Mongolia.

<i>Hagryphus</i> Extinct genus of dinosaurs

Hagryphus is a monospecific genus of caenagnathid dinosaur from southern Utah that lived during the Late Cretaceous in what is now the Kaiparowits Formation of the Grand Staircase–Escalante National Monument. The type and only species, Hagryphus giganteus, is known only from an incomplete but articulated left manus and the distal portion of the left radius. It was named in 2005 by Lindsay E. Zanno and Scott D. Sampson. Hagryphus has an estimated length of 2.4–3 metres and weight of 50 kilograms.

<i>Elmisaurus</i> Extinct genus of dinosaurs

Elmisaurus is an extinct genus of caenagnathid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was a theropod belonging to the Oviraptorosauria.

<i>Nemegtomaia</i> Extinct genus of dinosaurs

Nemegtomaia is a genus of oviraptorid dinosaur from what is now Mongolia that lived in the Late Cretaceous Period, about 70 million years ago. The first specimen was found in 1996, and became the basis of the new genus and species N. barsboldi in 2004. The original genus name was Nemegtia, but this was changed to Nemegtomaia in 2005, as the former name was preoccupied. The first part of the generic name refers to the Nemegt Basin, where the animal was found, and the second part means "good mother", in reference to the fact that oviraptorids are known to have brooded their eggs. The specific name honours the palaeontologist Rinchen Barsbold. Two more specimens were found in 2007, one of which was found on top of a nest with eggs, but the dinosaur had received its genus name before it was found associated with eggs.

<span class="mw-page-title-main">Caenagnathidae</span> Extinct family of dinosaurs

Caenagnathidae is a family of derived caenagnathoid dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids had specialized beaks, long necks, and short tails, and would have been covered in feathers. The relationships of caenagnathids were long a puzzle. The family was originally named by Raymond Martin Sternberg in 1940 as a family of flightless birds. The discovery of skeletons of the related oviraptorids revealed that they were in fact non-avian theropods, and the discovery of more complete caenagnathid remains revealed that Chirostenotes pergracilis, originally named on the basis of a pair of hands, and Citipes elegans, originally thought to be an ornithomimid, named from a foot, were caenagnathids as well.

<i>Gigantoraptor</i> Extinct genus of dinosaurs

Gigantoraptor is a genus of large oviraptorosaur dinosaur that lived in Asia during the Late Cretaceous period. It is known from the Iren Dabasu Formation of Inner Mongolia, where the first remains were found in 2005.

<i>Ganzhousaurus</i> Extinct genus of dinosaurs

Ganzhousaurus is an extinct genus of oviraptorid dinosaur known from the Late Cretaceous Nanxiong Formation of Nankang County, Ganzhou City of Jiangxi Province, southern China. It was found in a Maastrichtian deposit and contains a single species, Ganzhousaurus nankangensis. It is distinguished by a combination of primitive and derived features.

<i>Nankangia</i> Extinct genus of dinosaurs

Nankangia is an extinct genus of caenagnathoid oviraptorosaurian dinosaur known from the Upper Cretaceous Nanxiong Formation of Nankang County, Ganzhou City of Jiangxi Province, southeastern China. It contains a single species, Nankangia jiangxiensis. N. jiangxiensis coexisted with at least four other caenagnathoids, including but not limited to Corythoraptor, Banji, Ganzhousaurus and Jiangxisaurus. The relatively short dentary and non-downturned mandibular symphysis of Nankangia suggest that it may have been more herbivorous than carnivorous. Its diet consisted of leaves and seeds.

<i>Anzu wyliei</i> Genus of caenagnathid dinosaur

Anzu is a monospecific genus of caenagnathid dinosaur from North Dakota, South Dakota and Montana that lived during the Late Cretaceous in what is now the Hell Creek Formation. The type species and only species, Anzu wyliei is known from numerous skeletons that preserve cranial and postcranial elements. It was named in 2014 by Matthew C. Lamanna, Hans-Dieter Sues, Emma R. Schachner, and Tyler R. Lyson.

<span class="mw-page-title-main">Timeline of oviraptorosaur research</span>

This timeline of oviraptorosaur research is a chronological listing of events in the history of paleontology focused on the oviraptorosaurs, a group of beaked, bird-like theropod dinosaurs. The early history of oviraptorosaur paleontology is characterized by taxonomic confusion due to the unusual characteristics of these dinosaurs. When initially described in 1924 Oviraptor itself was thought to be a member of the Ornithomimidae, popularly known as the "ostrich" dinosaurs, because both taxa share toothless beaks. Early caenagnathid oviraptorosaur discoveries like Caenagnathus itself were also incorrectly classified at the time, having been misidentified as birds.

<i>Beibeilong</i> Caenagnathid dinosaur genus from the Late Cretaceous

Beibeilong is a genus of large caenagnathid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 96 million to 88 million years ago. The genus contains a single species, Beibeilong sinensis. The species was named and described in 2017 through analysis of an embryonic skeleton and partial nest with large eggs that were discovered in the Gaogou Formation of China between 1992 and 1993.

<i>Leptorhynchos gaddisi</i> Extinct species of dinosaur

Leptorhynchos is an extinct genus of caenagnathid theropod from the Late Cretaceous of what is now the US state of Texas, although it has been suggested to also exist in Alberta and South Dakota. The type species is L. gaddisi, and it is currently the only widely accepted valid species. The generic name of Leptorhynchos comes from the Greek "leptos" meaning "small" and "rhynchos" meaning "beak". The specific epithet is in honor of the Gaddis family, who owned the land on which the holotype was discovered.

References

  1. Xu, X., Tan, Q., Wang, J., Zhao, X., and Tan, L. (2007). "A gigantic bird-like dinosaur from the Late Cretaceous of China." Nature, 447: 844-847.
  2. 1 2 Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis, Indiana.
  3. Holtz, Thomas R. Jr. (2010) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2010 Appendix.
  4. Senter, P. (2007). "A new look at the phylogeny of Coelurosauria (Dinosauria: Theropoda)." Journal of Systematic Palaeontology, ( doi : 10.1017/S1477201907002143).

Further reading