Protarchaeopteryx Temporal range: Early Cretaceous, | |
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Holotype, Geological Museum of China | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | † Oviraptorosauria |
Genus: | † Protarchaeopteryx Ji & Ji, 1997 |
Species: | †P. robusta |
Binomial name | |
†Protarchaeopteryx robusta Ji & Ji, 1997 | |
Protarchaeopteryx (meaning "before Archaeopteryx " although it is from the Cretaceous and Archaeopteryx is from the Jurassic) is a genus of turkey-sized feathered theropod dinosaur from China. [1] Known from the Jianshangou bed of the Yixian Formation, it lived during the early Aptian age of the Early Cretaceous, approximately 124.6 million years ago. [2] It was probably a herbivore or omnivore, although its hands were very similar to those of small carnivorous dinosaurs. It appears to be one of the most basal members of the Oviraptorosauria, closely related to Incisivosaurus , [3] [4] or a taxon slightly less closely related to birds than oviraptorosaurs were. [5] [6]
The holotype and only known specimen of Protarchaeopteryx is NGMC 2125, a partial skeleton. [7]
Protarchaeopteryx had long legs, and could have been a quick runner. It had well-developed, vaned feathers extended from a relatively short tail; the hands were long and slender, and had three fingers with sharp, curved claws. Its bones were hollow and bird-like, and it possessed a wishbone. [8] At around 1 metre (3.3 ft) in length, it would have been larger than Archaeopteryx. [8] In 2016, Gregory S. Paul gave a lower estimate of 0.7 metres (2.3 ft) in length and 1.6 kilograms (3.5 lb) in body mass. [9] Protarchaeopteryx also had symmetrical feathers on its tail. Since modern birds that have symmetrical feathers are flightless, and the skeletal structure of Protarchaeopteryx would not support flapping flight, it is assumed that it was flightless as well. [10] It has been suggested that it could have had an arboreal lifestyle, jumping from tree limbs and using its forelimbs for a form of parachuting. [11]
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens, have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like Sinovenator, demonstrate striking anatomical similarities with Archaeopteryx and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves.
Deinonychosauria is a clade of paravian dinosaurs which lived from the Late Jurassic to the Late Cretaceous periods. Fossils have been found across the globe in North America, Europe, Africa, Asia, South America, and Antarctica, with fossilized teeth giving credence to the possibility that they inhabited Australia as well. This group of dinosaurs are known for their sickle-shaped toe claws and features in the shoulder bones.
Caudipteryx is a genus of small oviraptorosaur dinosaurs that lived in Asia during the Early Cretaceous, around 124.6 million years ago. They were feathered and extremely birdlike in their overall appearance, to the point that some paleontologists suggested it was a bird. Two species have been described: C. zoui, in 1998, and C. dongi, in 2000.
Oviraptoridae is a group of bird-like, herbivorous and omnivorous maniraptoran dinosaurs. Oviraptorids are characterized by their toothless, parrot-like beaks and, in some cases, elaborate crests. They were generally small, measuring between one and two metres long in most cases, though some possible oviraptorids were enormous. Oviraptorids are currently known only from the Late Cretaceous of Asia, with the most well-known species and complete specimens found only in the Gobi Desert of Mongolia and northwestern China.
Oviraptorosaurs are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from Caudipteryx, which was the size of a turkey, to the 8-meter-long, 1.4-ton Gigantoraptor. The group is close to the ancestry of birds. Some researchers such as Maryanska et al (2002) and Osmólska et al. (2004) have proposed that they may represent primitive flightless birds. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.
Jinfengopteryx is a genus of maniraptoran dinosaur. It was found in the Qiaotou Member of the Huajiying Formation of Hebei Province, China, and is therefore of uncertain age. The Qiaotou Member may correlate with the more well-known Early Cretaceous Yixian Formation, and so probably dates to around 122 Ma ago.
Yixianosaurus is a maniraptoran theropod dinosaur genus from the Early Cretaceous of China.
Alxasaurus is a genus of therizinosauroid theropod dinosaurs from the Early Cretaceous Bayin-Gobi Formation of Inner Mongolia.
Jixiangornis is a genus of basal avialan dinosaurs from the Early Cretaceous. Like later avialans, it had no teeth, but it also had a long tail, unlike modern birds. Since teeth were still present in some more derived short-tailed avialans, Jixiangornis seems to have evolved its toothlessness independently of modern birds. The long forelimb indicates at least some aerial ability. Jixiangornis is currently known only from a single specimen, a complete but juvenile skeleton. The fossil was found in the Yixian Formation near Beipiao City, western Liaoning, China.
Archaeopterygidae is a group of paravian dinosaurs, known from the latest Jurassic and earliest Cretaceous of Europe. In most current classifications, it contains only the genera Archaeopteryx and Wellnhoferia. As its name suggests, Protarchaeopteryx was also once referred to this group, but most paleontologists now consider it an oviraptorosaur. Other referred genera, like Jurapteryx, Wellnhoferia, and "Proornis", are probably synonymous with Archaeopteryx or do not belong into this group. Jinfengopteryx was originally described as an archaeopterygid, though it was later shown to be a troodontid. A few studies have recovered Anchiornis and Xiaotingia to also be members of the Archaeopterygidae, though most subsequent analyses have failed to arrive at the same result. Uncertainties still exist, however, and it may not be possible to confidently state whether archaeopterygids are more closely related to modern birds or to deinonychosaurs barring new and better specimens of relevant species. Teeth attributable to archaeopterygids are known from the earliest Cretaceous (Berriasian) Cherves-de-Cognac locality and the Angeac-Charente bonebed of France.
The scientific question of within which larger group of animals birds evolved has traditionally been called the "origin of birds". The present scientific consensus is that birds are a group of maniraptoran theropod dinosaurs that originated during the Mesozoic era.
Compsognathidae is a family of coelurosaurian theropod dinosaurs. Compsognathids were small carnivores, generally conservative in form, hailing from the Jurassic and Cretaceous Periods. The bird-like features of these species, along with other dinosaurs such as Archaeopteryx inspired the idea for the connection between dinosaur reptiles and modern-day avian species. Compsognathid fossils preserve diverse integument — skin impressions are known from four genera commonly placed in the group, Compsognathus, Sinosauropteryx, Sinocalliopteryx and Juravenator. While the latter three show evidence of a covering of some of the earliest primitive feathers over much of the body, Juravenator and Compsognathus also show evidence of scales on the tail or hind legs. "Ubirajara jubatus", informally described in 2020, had elaborate integumentary structures on its back and shoulders superficially similar to the display feathers of a standardwing bird-of-paradise, and unlike any other non-avian dinosaur currently described.
Avialae is a clade containing the only living dinosaurs, the birds, and their closest relatives. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used.
Zhongornis is a genus of primitive maniraptoran dinosaurs that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008.
Hesperonychus is a genus of small paravian theropod dinosaur. It may be a dromaeosaurid or an avialan. There is one described species, Hesperonychus elizabethae. The type species was named in honor of Dr. Elizabeth Nicholls of the Royal Tyrrell Museum of Palaeontology who collected it as a student in 1982. It is known from fossils recovered from the Dinosaur Park Formation and possibly from the uppermost strata of the Oldman Formation of Alberta, dating to the Campanian stage of the Late Cretaceous around 75 million years ago.
Xiaotingia is a genus of anchiornithid theropod dinosaur from Middle Jurassic or early Late Jurassic deposits of western Liaoning, China, containing a single species, Xiaotingia zhengi.
Ningyuansaurus is a pennaraptoran dinosaur genus. It contains the single species Ningyuansaurus wangi, known from a fossil specimen from the Early Cretaceous Yixian Formation of Jianchang, western Liaoning Province, People's Republic of China. It was originally thought to be the basalmost species of oviraptorosaur, based on its long skull and a greater number of teeth in comparison to any other known oviraptorosaur. However, later research showed that it is a member of Halszkaraptorinae. The generic name Ningyuansaurus is derived from Ningyuan, an ancient name for Xingcheng City. The specific name honors Wang Qiuwu, the private owner of the specimen who donated it for scientific study. The specimen is now in the Confuciusornis Museum in Xingcheng.
Anchiornithidae is a family of small paravian dinosaurs. Anchiornithids have been classified at varying positions in the paravian tree, with some scientists classifying them as a distinct family, a basal subfamily of Troodontidae, members of Archaeopterygidae, or an assemblage of dinosaurs that are an evolutionary grade within Avialae or Paraves.