| Borogovia Temporal range: Late Cretaceous, | |
|---|---|
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| Leg bones of the holotype | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Dinosauria |
| Clade: | Saurischia |
| Clade: | Theropoda |
| Family: | † Troodontidae |
| Genus: | † Borogovia Osmólska, 1987 |
| Type species | |
| †Borogovia gracilicrus Osmólska, 1987 | |
Borogovia is a troodontid theropod dinosaur genus which lived during the Late Cretaceous Period, in what is now Mongolia. It contains one species, Borogovia gracilicrus.
In 1971, a Polish-Mongolian expedition discovered the remains of a small theropod at the Altan Uul IV site, in the Nemegt Basin of Ömnögovĭ province. In 1982, upon describing Hulsanpes perlei , the find was reported by Halszka Osmólska and considered by her to be a possible specimen of Saurornithoides . [1] Later she concluded that it represented a species new to science.
In 1987, Osmólska named and described the type species Borogovia gracilicrus. The generic name is derived from the fantasy creatures known as 'borogoves' in the Lewis Carroll poem "Jabberwocky" in Through the Looking-Glass . The specific name is a combination of Latin gracilis, "lightly built", and crus, "shin", in reference to the elegant build of the lower leg. [2]
The holotype specimen, ZPAL MgD-I/174, was found in the Nemegt Formation, dating from the early Maastrichtian. It consists of two lower legs of a single individual, including fragments of both tibiotarsi, the undersides of both metatarsi, and the second, third, and fourth toes of each foot. [2]
The tibiotarsi have an estimated length of twenty-eight centimetres. Borogovia is about two meters (6 feet) long, weighing some twenty kilograms (forty-five pounds). The tibiotarsus is very elongated. The third toe is narrow. The second phalanx of the second toe is short. The claw of the second toe is short and relatively flat. Osmólska claimed that the second toe could not be hyperextended and suggested that it had regained a weight-bearing function, compensating for the weakness of the third toe. [2] In 2021, Andrea Cau and Daniel Madzia have called this the "falchiporan condition". [3]
Osmólska (1987) assigned Borogovia to the Troodontidae in 1987, noting that it may be a junior synonym of Saurornithoides. [2] In the 2021 description of Tamarro , Sellés et al. recovered Borogovia as a member of the Troodontinae, closely related to Saurornithoides. [4] Later in 2021, Cau and Madzia revised material of Borogovia and recovered it as a sister taxon to Troodontinae. [3]
Sellés et al., 2021 [4]
| Cau & Madzia, 2021 [3] |
Troodontidae is a clade of bird-like theropod dinosaurs from the Late Jurassic to Late Cretaceous. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens, have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like Sinovenator, demonstrate striking anatomical similarities with Archaeopteryx and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves.
Avimimus, meaning "bird mimic", is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 to 70 million years ago.
Saurornithoides is a genus of troodontid maniraptoran dinosaur, which lived during the Late Cretaceous period. These creatures were predators, which could run fast on their hind legs and had excellent sight and hearing. The name is derived from the Greek stems saur~ (lizard), ornith~ (bird) and eides (form), referring to its bird-like skull.
Adasaurus is a genus of dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous period about 70 million years ago. The genus is known from two partial specimens found in the Nemegt Formation of Mongolia that were partially described in 1983 by the paleontologist Rinchen Barsbold.
Tylocephale is a genus of pachycephalosaurid dinosaur, a group of dome-headed, herbivorous ornithischians, that lived during the Late Campanian stage of the Late Cretaceous in what is now Mongolia. It is known from a partial skull and associated mandible that were unearthed in 1971 by a Polish-Mongolian Expedition to the Barun Goyot Formation of the Gobi Desert. The specimen was described in 1974 by Polish paleontologists Teresa Maryańska and Halszka Osmólska as a new genus and species.
Tochisaurus is a genus of small troodontid theropod dinosaur from the Late Cretaceous Period of Mongolia. The type species is Tochisaurus nemegtensis.
Hulsanpes is a genus of halszkaraptorine theropod dinosaurs that lived during the Late Cretaceous in what is now the Barun Goyot Formation of Mongolia, about 75-72 million years ago. The remains were found in 1970 and formally described in 1982 by Halszka Osmólska, who noted that the genus is represented by an immature individual. Hulsanpes represents the first record of the basal dromaeosaurid subfamily Halszkaraptorinae.
The Barun Goyot Formation is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia.
The Nemegt Formation is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds.
Halszka Osmólska was a Polish paleontologist who had specialized in Mongolian dinosaurs.
Mahakala is a genus of halszkaraptorine theropod dinosaur from the Campanian-age Upper Cretaceous Djadokhta Formation of Ömnögovi, Mongolia. It is based on a partial skeleton found in the Gobi Desert. Mahakala was a small dromaeosaurid, and its skeleton shows features that are also found in early troodontids and avialans. Despite its late appearance, it is among the most basal dromaeosaurids. Its small size, and the small size of other basal deinonychosaurians, suggests that small size appeared before flight capability in birds. The genus is named for Mahakala, one of eight protector deities (dharmapalas) in Tibetan Buddhism.
Xixiasaurus is a genus of troodontid dinosaur that lived during the Late Cretaceous Period in what is now China. The only known specimen was discovered in Xixia County, Henan Province, in central China, and became the holotype of the new genus and species Xixiasaurus henanensis in 2010. The names refer to the areas of discovery, and can be translated as "Henan Xixia lizard". The specimen consists of an almost complete skull, part of the lower jaw, and teeth, as well as a partial right forelimb.
Gobivenator is an extinct genus of troodontid theropod dinosaur known from the late Campanian Djadokhta Formation of central Gobi Desert, Mongolia. It contains a single species, Gobivenator mongoliensis. G. mongoliensis is known from a single individual, which represents the most complete specimen of a Late Cretaceous troodontid currently known.
This timeline of troodontid research is a chronological listing of events in the history of paleontology focused on the troodontids, a group of bird-like theropod dinosaurs including animals like Troodon. Troodontid remains were among the first dinosaur fossils to be reported from North America after paleontologists began performing research on the continent, specifically the genus Troodon itself. Since the type specimen of this genus was only a tooth and Troodon teeth are unusually similar to those of the unrelated thick-headed pachycephalosaurs, Troodon and its relatives would be embroiled in taxonomic confusion for over a century. Troodon was finally recognized as distinct from the pachycephalosaurs by Phil Currie in 1987. By that time many other species now recognized as troodontid had been discovered but had been classified in the family Saurornithoididae. Since these families were the same but the Troodontidae named first, it carries scientific legitimacy.
Halszkaraptorinae is an extinct basal ("primitive") subfamily of Dromaeosauridae that includes the enigmatic genera Halszkaraptor, Natovenator, Mahakala, and Hulsanpes. Halszkaraptorines are definitively known only from Late Cretaceous strata in Asia, specifically in Mongolia. It is debated among researchers whether the group had a semiaquatic lifestyle.
Khulsanurus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Barungoyot Formation of the Khulsan Locality in the Gobi Desert region of Mongolia. The type and only species is Khulsanurus magnificus.
Ondogurvel is a genus of alvarezsaurid dinosaur from the Late Cretaceous (Campanian) Barun Goyot Formation in southern Mongolia. The type and only species is O. alifanovi, known from a partial skeleton consisting of fragments of two last dorsal vertebrae, three anterior sacral vertebrae, right ilium, left and right pubis and ischium, articulated right tibia, fibula, metatarsals II and IV, and phalanges IV-1 and IV-2, right carpometacarpus, left and right manual phalanx II-1, right femur, left pedal phalanx II-1, and fragments of unidentified phalanges.
Natovenator is a genus of halszkaraptorine dromaeosaurid dinosaur from the Late Cretaceous Barun Goyot Formation of Mongolia. The genus is known from a single species, N. polydontus. Natovenator is crucial to the understanding of halszkaraptorines due to it providing more support for the semi-aquatic lifestyle that has been proposed for this clade. This discovery is important as the semi-aquatic lifestyles of halszkaraptorines was contested in early 2022.
Jaculinykus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Baruungoyot Formation of Mongolia. The genus contains a single species, J. yaruui, known from a nearly complete articulated skeleton including bones of the skull. Jaculinykus is notable for its unique hand, which has a hypertrophied first digit and greatly reduced second digit, which is intermediate between the tridactyl hand of Shuvuuia and monodactyl hand of Linhenykus.
Harenadraco is an extinct genus of troodontid theropod dinosaurs from the Late Cretaceous Baruungoyot Formation of Mongolia. The genus contains a single species, H. prima, known from a fragmentary skeleton. Harenadraco represents the first troodontid known from the Baruungoyot Formation. Since members of this family are also known from the Nemegt and Djadochta Formations, the discovery of Harendraco indicates that troodontids were important faunal components of every formation within the Nemegt Basin.
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