Zanabazar junior

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Zanabazar junior
Temporal range: Late Cretaceous,
~70  Ma
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Zanabazar skeletal.png
Skeletal restoration of IGM 100/1
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: Troodontidae
Subfamily: Troodontinae
Genus: Zanabazar
Norell et al., 2009
Species:
Z. junior
Binomial name
Zanabazar junior
Synonyms

Zanabazar is a genus of large troodontid dinosaurs from the Late Cretaceous of Mongolia. The genus was originally named by Rinchen Barsbold as the new species Saurornithoides junior. In 2009 it was reclassified as its own genus and species, Zanabazar junior, named after Zanabazar, the first spiritual figurehead of Tibetan Buddhism in Mongolia. [1] The holotype includes a skull, vertebrae, and right hindlimb. Zanabazar was one of the largest and most derived troodontids.

Contents

History of discovery

Reconstructed skull of IGM 100/1 Zanabazar junior Headden.jpg
Reconstructed skull of IGM 100/1

The holotype was discovered in 1964 from the Bügiin Tsav locality of the Nemegt Formation and initially described by Rinchen Barsbold as a new species of Saurornithoides (S. junior) in 1974. This specimen, IGM 100/1, includes a nearly complete skull and braincase, part of the pelvis, some tail vertebrae, and parts of the right hindlimb. [2] In 2009 a review of the genus found that the support for S. junior in the same genus as S. mongoliensis was lacking. Mark Norell and colleagues re-classified the species in the new genus Zanabazar, which they named in honor of Zanabazar, the first spiritual head (Bogd Gegen) of Tibetan Buddhism in Outer Mongolia. [1]

Description

Size comparison of the holotype Zanabazar Scale.svg
Size comparison of the holotype

Zanabazar were large troodontids reaching 2.3 m (7.5 ft) in length and weighing 25 kg (55 lb). [3] They are the largest known Asian troodontids, with a skull length of 27.2 cm (272 mm). At the time of the discovery of the genus, the only other troodontids that appeared to be larger than it were specimens from Alaska, [1] however, Latenivenatrix are now considered the largest troodontids with 3.5 m (11 ft) in length. [4] The preserved vertebrae in IGM 100/1 are completely fused, indicating that this individual was an adult at the time of death. [1]

Classification

While originally included in Saurornithoides, within the family Saurornithoididae, [2] Zanabazar is now thought to be a derived member of Troodontidae. [1]

Life restoration Zanabazar.jpg
Life restoration
Comparison of troodontid teeth; C, D, F, and G are Zanabazar Troodontid teeth.jpg
Comparison of troodontid teeth; C, D, F, and G are Zanabazar

The cladogram below shows the phylogenetic position of Zanabazar among other troodontids following a 2014 analysis. [5]

Paraves  

Avialae

  Deinonychosauria  

Dromaeosauridae

  Troodontidae  

Sinovenator

Anchiornis

Mei

Talos

Byronosaurus

IGM 100/44

Sinornithoides

Linhevenator

Philovenator

Gobivenator

Troodon

Saurornithoides

Zanabazar

See also

Related Research Articles

<span class="mw-page-title-main">Troodontidae</span> Extinct family of bird-like dinosaurs

Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens, have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like Sinovenator, demonstrate striking anatomical similarities with Archaeopteryx and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves.

<i>Troodon</i> Extinct genus of dinosaurs

Troodon is a former wastebasket taxon and a potentially dubious genus of relatively small, bird-like theropod dinosaurs definitively known from the Campanian age of the Late Cretaceous period. It includes at least one species, Troodon formosus, known from Montana. Discovered in October 1855, T. formosus was among the first dinosaurs found in North America, although it was thought to be a lizard until 1877. Several well-known troodontid specimens from the Dinosaur Park Formation in Alberta were once believed to be members of this genus. However, recent analyses in 2017 have found this genus to be undiagnostic and referred some of these specimens to the genus Stenonychosaurus some to the genus Latenivenatrix, and some to the genus Pectinodon. The genus name is Ancient Greek for "wounding tooth", referring to the teeth, which were different from those of most other theropods known at the time of their discovery. The teeth bear prominent, apically oriented serrations. These "wounding" serrations, however, are morphometrically more similar to those of herbivorous reptiles, and suggest a possibly omnivorous diet.

<i>Oviraptor</i> Extinct genus of dinosaurs

Oviraptor is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period. The first remains were collected from the Djadokhta Formation of Mongolia in 1923 during a paleontological expedition led by Roy Chapman Andrews, and in the following year the genus and type species Oviraptor philoceratops were named by Henry Fairfield Osborn. The genus name refers to the initial thought of egg-stealing habits, and the specific name was intended to reinforce this view indicating a preference over ceratopsian eggs. Despite the fact that numerous specimens have been referred to the genus, Oviraptor is only known from a single partial skeleton regarded as the holotype, as well as a nest of about fifteen eggs and several small fragments from a juvenile.

<i>Anserimimus</i> Extinct genus of dinosaurs

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<i>Enigmosaurus</i> Extinct genus of reptiles

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<i>Saurornithoides</i> Extinct genus of dinosaurs

Saurornithoides is a genus of troodontid maniraptoran dinosaur, which lived during the Late Cretaceous period. These creatures were predators, which could run fast on their hind legs and had excellent sight and hearing. The name is derived from the Greek stems saur~ (lizard), ornith~ (bird) and eides (form), referring to its bird-like skull.

<i>Byronosaurus</i> Extinct genus of dinosaurs

Byronosaurus is a genus of troodontid dinosaur from the Late Cretaceous Period of Mongolia.

<i>Adasaurus</i> Extinct genus of dinosaurs

Adasaurus is a genus of dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous period about 70 million years ago. The genus is known from two partial specimens found in the Nemegt Formation of Mongolia that were partially described in 1983 by the paleontologist Rinchen Barsbold.

<i>Sinornithoides</i> Extinct genus of dinosaurs

Sinornithoides is a genus of troodontid theropod dinosaurs containing the single species Sinornithoides youngi. S. youngi lived during the Early Cretaceous. It measured approximately one meter long (3.3 ft). It lived in Inner Mongolia, China, and probably ate invertebrates and other small prey. They lived in what is now Mongolia, which was part of Laurasia.

<i>Mononykus</i> Extinct genus of dinosaurs

Mononykus is a genus of alvarezsaurid dinosaur that lived during the Late Cretaceous in what is now Asia on the Nemegt Formation, about 70 million years ago.

<i>Citipati</i> Genus of oviraptorid dinosaur

Citipati is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality at the Djadochta Formation, where the first remains were collected during the 1990s. The genus and type species Citipati osmolskae were named and described in 2001. A second species from the adjacent Zamyn Khondt locality may also exist. Citipati is one of the best-known oviraptorids thanks to a number of well-preserved specimens, including individuals found in brooding positions atop nests of eggs, though most of them were initially referred to the related Oviraptor. These nesting specimens have helped to solidify the link between non-avian dinosaurs and birds.

<i>Sinovenator</i> Extinct genus of dinosaurs

Sinovenator is a genus of troodontid dinosaur from China. It is from the early Cretaceous Period.

<i>Sinusonasus</i> Extinct genus of dinosaurs

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<i>Tsaagan</i> Extinct genus of dinosaurs

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<span class="mw-page-title-main">Mahakala omnogovae</span> Extinct species of dinosaur

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<i>Linhevenator</i> Extinct genus of dinosaurs

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<i>Gobivenator</i> Extinct genus of dinosaurs

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<span class="mw-page-title-main">Timeline of troodontid research</span> Events in the history of paleontology

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<i>Liaoningvenator</i> Extinct genus of dinosaurs

Liaoningvenator is a genus of troodontid theropod dinosaur from the Early Cretaceous of China. It contains a single species, L. curriei, named after paleontologist Phillip J. Currie in 2017 by Shen Cai-Zhi and colleagues from an articulated, nearly complete skeleton, one of the most complete troodontid specimens known. Shen and colleagues found indicative traits that placed Liaoningvenator within the Troodontidae. These traits included its numerous, small, and closely packed teeth, as well as the vertebrae towards the end of its tail having shallow grooves in place of neural spines on their top surfaces.

<i>Almas ukhaa</i> Genus of reptiles (fossil)

Almas is a genus of troodontid theropod dinosaur from the Late Cretaceous of Mongolia. It contains a single species, Almas ukhaa, named in 2017 by Pei Rui and colleagues, based on a partial articulated skeleton. The only known specimen was found in the Djadochta Formation, which is late Campanian in age.

References

  1. 1 2 3 4 5 Norell, M. A.; Makovicky, P. J.; Bever, G. S.; Balanoff, A. M.; Clark, J. M.; Barsbold, R.; Rowe, T. (2009). "A review of the Mongolian Cretaceous dinosaur Saurornithoides (Troodontidae, Theropoda)". American Museum Novitates (3654): 1−63. doi:10.1206/648.1. hdl: 2246/5973 .
  2. 1 2 Barsbold, R. (1974). "Saurornithoididae, a new family of small theropod dinosaurs from Central Asia and North America" (PDF). Palaeontologica Polonica. 30: 5−22.
  3. Paul, G. S. (2016). The Princeton Field Guide to Dinosaurs (2nd ed.). Princeton, New Jersey: Princeton University Press. p. 161. ISBN   9780691167664.
  4. van der Reest, A. J.; Currie, P. J. (2017). "Troodontids (Theropoda) from the Dinosaur Park Formation, Alberta, with a description of a unique new taxon: implications for deinonychosaur diversity in North America". Canadian Journal of Earth Sciences. 54 (9): 919−935. Bibcode:2017CaJES..54..919V. doi:10.1139/cjes-2017-0031. hdl: 1807/78296 . ISSN   0008-4077.
  5. Tsuihiji, T.; Barsbold, R.; Watabe, M.; Tsogtbaatar, K.; Chinzorig, T.; Fujiyama, Y.; Suzuki, S. (2014). "An exquisitely preserved troodontid theropod with new information on the palatal structure from the Upper Cretaceous of Mongolia". Naturwissenschaften. 101 (2): 131−142. Bibcode:2014NW....101..131T. doi:10.1007/s00114-014-1143-9. PMID   24441791.