Archaeopterygidae Possible | |
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Berlin specimen of Archaeopteryx lithographica | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Paraves |
Family: | † Archaeopterygidae Huxley, 1871 (conserved name) |
Type species | |
† Archaeopteryx lithographica von Meyer, 1861 (conserved name) | |
Genera and possible subfamily [1] | |
Synonyms | |
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Archaeopterygidae is a group of paravian dinosaurs, known from the latest Jurassic and earliest Cretaceous of Europe. In most current classifications, it contains only the genera Archaeopteryx and Wellnhoferia . As its name suggests, Protarchaeopteryx was also once referred to this group, but most paleontologists now consider it an oviraptorosaur. Other referred genera, like Jurapteryx , Wellnhoferia , and "Proornis", are probably synonymous with Archaeopteryx (the former two) or do not belong into this group (the last). Jinfengopteryx was originally described as an archaeopterygid, though it was later shown to be a troodontid. [3] [4] [5] A few studies have recovered Anchiornis and Xiaotingia (usually considered part of a distinct clade, Anchiornithidae) to also be members of the Archaeopterygidae, [6] though most subsequent analyses have failed to arrive at the same result. Uncertainties still exist, however, and it may not be possible to confidently state whether archaeopterygids are more closely related to modern birds or to deinonychosaurs barring new and better specimens of relevant species. [7] Teeth attributable to archaeopterygids are known from the earliest Cretaceous (Berriasian) Cherves-de-Cognac locality and the Angeac-Charente bonebed of France. [8] [9]
The family Archaeopterygidae is the only family in the order Archaeopterygiformes, which was coined by Max Fürbringer in 1888 to contain Archaeopterygidae and genus Archaeopteryx. [10] A formal phylogenetic definition for Archaeopterygidae was given by Xu and colleagues in 2011: the clade comprising all animals closer to Archaeopteryx than to the house sparrow or Dromaeosaurus . [6]
The family Dromaeosauridae, traditionally considered to be non-avian dinosaurs, have been included in this group by at least one author, although the group was paraphyletic in that classification, with Dromaeosaurus and Velociraptor (including Deinonychus and Saurornitholestes ) being more closely related to modern birds than Archaeopteryx was. [11] Discoveries of a number of primitive forms have muddied the relationships of early birds, making it possible that Velociraptor and Deinonychus could be considered birds as they might have evolved from flying ancestors. Palaeoartist Gregory S. Paul placed dromaeosaurids in Archaeopterygidae for these reasons, though the eventual cladistic definition of Archaeopterygidae explicitly excluded them. [11]
The family Anchiornithidae has had some of the members or the entirety of the group placed as archaeopterygids in various systematic studies. [6] [12] [1] The cladogram below shows the results of the phylogenetic analysis by Cau (2020). [1]
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Velociraptor is a genus of small dromaeosaurid dinosaurs that lived in Asia during the Late Cretaceous epoch, about 75 million to 71 million years ago. Two species are currently recognized, although others have been assigned in the past. The type species is V. mongoliensis, named and described in 1924. Fossils of this species have been discovered in the Djadochta Formation, Mongolia. A second species, V. osmolskae, was named in 2008 for skull material from the Bayan Mandahu Formation, China.
Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discoveries of complete and articulated specimens, have helped to increase understanding about this group. Anatomical studies, particularly studies of the most primitive troodontids, like Sinovenator, demonstrate striking anatomical similarities with Archaeopteryx and primitive dromaeosaurids, and demonstrate that they are relatives comprising a clade called Paraves.
Utahraptor is a genus of large dromaeosaurid dinosaur that lived during the Early Cretaceous period from around 135 to 130 million years ago in what is now the United States. The genus was described in 1993 by an American paleontologist James Kirkland and colleagues with the type species Utahraptor ostrommaysi, based on fossils that had been unearthed earlier from the Cedar Mountain Formation of Utah. Later, many additional specimens were described including those from the skull and postcranium in addition to those of younger individuals.
Deinonychosauria is a clade of paravian dinosaurs which lived from the Late Jurassic to the Late Cretaceous periods. Fossils have been found across the globe in North America, Europe, Africa, Asia, South America, and Antarctica, with fossilized teeth giving credence to the possibility that they inhabited Australia as well. This group of dinosaurs are known for their sickle-shaped toe claws and features in the shoulder bones.
Dromaeosauridae is a family of feathered coelurosaurian theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek δρομαῖος (dromaîos), meaning 'running at full speed', 'swift', and σαῦρος (saûros), meaning 'lizard'. In informal usage, they are often called raptors, a term popularized by the film Jurassic Park; several genera include the term "raptor" directly in their name, and popular culture has come to emphasize their bird-like appearance and speculated bird-like behavior.
Protarchaeopteryx is a genus of turkey-sized feathered theropod dinosaur from China. Known from the Jianshangou bed of the Yixian Formation, it lived during the early Aptian age of the Early Cretaceous, approximately 124.6 million years ago. It was probably a herbivore or omnivore, although its hands were very similar to those of small carnivorous dinosaurs. It appears to be one of the most basal members of the Oviraptorosauria, closely related to Incisivosaurus, or a taxon slightly less closely related to birds than oviraptorosaurs were.
Pedopenna is a genus of small, feathered, maniraptoran dinosaur from the Daohugou Beds in China. It is possibly older than Archaeopteryx, though the age of the Daohugou Beds where it was found is debated. A majority of studies suggest that beds probably date from between the late Middle Jurassic and early Late Jurassic Period.
Buitreraptor is a genus of dromaeosaurid dinosaurs that lived during the Late Cretaceous of Argentina at the Candeleros Formation. Buitreraptor was described in 2005 and the type species is Buitreraptor gonzalezorum. It was rooster-sized and had a very elongated head with many small teeth.
Adasaurus is a genus of dromaeosaurid dinosaur that lived in Asia during the Late Cretaceous period about 70 million years ago. The genus is known from two partial specimens found in the Nemegt Formation of Mongolia that were partially described in 1983 by the paleontologist Rinchen Barsbold.
Jinfengopteryx is a genus of maniraptoran dinosaur. It was found in the Qiaotou Member of the Huajiying Formation of Hebei Province, China, and is therefore of uncertain age. The Qiaotou Member may correlate with the more well-known Early Cretaceous Yixian Formation, and so probably dates to around 122 Ma ago.
Rahonavis is a genus of bird-like theropods from the Late Cretaceous of what is now northwestern Madagascar. It is known from a partial skeleton found by Catherine Forster and colleagues in Maevarano Formation rocks at a quarry near Berivotra, Mahajanga Province. Rahonavis was a small predator, at about 70 centimetres (2.3 ft) long and 0.45-2.27 kg, with the typical dromaesaurid-like raised sickle claw on the second toe. It was originally the first African coelurosaur until the discovery of Nqwebasaurus in 2000.
Jixiangornis is a genus of basal avialan dinosaurs from the Early Cretaceous. Like later avialans, it had no teeth, but it also had a long tail, unlike modern birds. Since teeth were still present in some more derived short-tailed avialans, Jixiangornis seems to have evolved its toothlessness independently of modern birds. The long forelimb indicates at least some aerial ability. Jixiangornis is currently known only from a single specimen, a complete but juvenile skeleton. The fossil was found in the Yixian Formation near Beipiao City, western Liaoning, China.
Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. The earliest velociraptorines are probably Nuthetes from the United Kingdom, and possibly Deinonychus from North America. However, several indeterminate velociraptorines have also been discovered, dating to the Kimmeridgian stage, in the Late Jurassic Period. These fossils were discovered in the Langenberg quarry, Oker near Goslar, Germany.
Avialae is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used.
Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avialans, which include diverse extinct taxa as well as the over 10,000 species of living birds. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research.
Mahakala is a genus of halszkaraptorine theropod dinosaur from the Campanian-age Upper Cretaceous Djadokhta Formation of Ömnögovi, Mongolia. It is based on a partial skeleton found in the Gobi Desert. Mahakala was a small dromaeosaurid, and its skeleton shows features that are also found in early troodontids and avialans. Despite its late appearance, it is among the most basal dromaeosaurids. Its small size, and the small size of other basal deinonychosaurians, suggests that small size appeared before flight capability in birds. The genus is named for Mahakala, one of eight protector deities (dharmapalas) in Tibetan Buddhism.
Balaur is a genus of theropod dinosaur from the late Cretaceous period, in what is now Romania. It is the type species of the monotypic genus Balaur, after the balaur, a dragon of Romanian folklore. The specific name bondoc means "stocky", so Balaur bondoc means "stocky dragon" in Romanian. This name refers to the greater musculature that Balaur had compared to its relatives. The genus, which was first described by scientists in August 2010, is known from two partial skeletons.
Xiaotingia is a genus of anchiornithid theropod dinosaur from Middle Jurassic or early Late Jurassic deposits of western Liaoning, China, containing a single species, Xiaotingia zhengi.
Jinfengopteryginae is a subfamily of bird-like theropod dinosaurs known from the Cretaceous of Eurasia. This group includes relatively few genera, with members discovered in 2005 but the name erected in 2012. Like other troodontids, this group of dinosaurs resided in the Paraves potentially close to the Avialae.
Anchiornithidae is a family of small paravian dinosaurs. Anchiornithids have been classified at varying positions in the paravian tree, with some scientists classifying them as a distinct family, a basal subfamily of Troodontidae, members of Archaeopterygidae, or an assemblage of dinosaurs that are an evolutionary grade within Avialae or Paraves.
Catalogue of fossil birds