Alvarezsaurus

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Alvarezsaurus
Temporal range: Late Cretaceous, 86–83  Ma
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Alvarezsaurus.jpg
Skeleton restoration
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: Alvarezsauridae
Genus: Alvarezsaurus
Bonaparte, 1991
Species:
A. calvoi
Binomial name
Alvarezsaurus calvoi
Bonaparte, 1991
Synonyms

Alvarezsaurus ( /ˌælvərɛzˈsɔːrəs/ ; "Alvarez's lizard") is a genus of alvarezsaurid dinosaur from the Late Cretaceous, living in Argentina approximately 86 - 83 million years ago. It was a small dinosaur, measuring 1–1.4 m (3.3–4.6 ft) long and weighing approximately 3 kg (6.6 lb). [1] [2] It was found in the Bajo de la Carpa Formation and was named by paleontologist José Bonaparte in 1991 after the historian Don Gregorio Alvarez. [3] The type species is A. calvoi.

Description

Restoration. Alvarezsaurus calvoi.jpg
Restoration.

Alvarezsaurus was a bipedal theropod. Like other lightweight theropods, it had a long tail, and its leg structure suggests that it was a fast runner. The most proximal elements of Alvarezsaurus caudal vertebrae exhibited ventrally sharp centra and the transverse processes of these vertebrae were sub-triangular and laterodistally directed, features seen in other alvarezsaurids like Shuvuuia. [4] Spinal processes were entirely absent or poorly developed, and each caudal vertebra supported short prezygapophyses. The scapula was visibly curved and proportionally smaller than those of other alvarezsaurids, and unlike its relatives Alvarezsaurus did not have a fused astragalus and calcaneum. It was unique in that its metatarsal III was its longest, followed by an unusually long metatarsal IV. It may have been insectivorous.[ citation needed ]

Classification

Skeletal diagram of known remains. Alvarezsaurus Jaime Headden.jpg
Skeletal diagram of known remains.

Alvarezsaurus is considered basal to better-known members of its family, such as Mononykus and Shuvuuia . It has been alternately classified with both non-avian theropod dinosaurs and early birds, but a move of the alvarezsaurids to be recognized as more closely related to neornithine birds proved controversial despite being supported by earlier studies. [4] It was once believed that the Patagonian alvarezsaur taxa were the most basal of their family, but the discovery of a more basal member, Haplocheirus , disproved that when its fossils were discovered in China. [5]

Related Research Articles

<i>Shuvuuia</i> Extinct family of bird-like dinosaurs

Shuvuuia is a genus of bird-like theropod dinosaur from the late Cretaceous period of Mongolia. It is a member of the family Alvarezsauridae, small coelurosaurian dinosaurs which are characterized by short but powerful forelimbs specialized for digging. The type species is Shuvuuia deserti, or "desert bird". The name Shuvuuia is derived from the Mongolian word shuvuu (шувуу) meaning "bird".

<i>Parvicursor</i> Extinct genus of dinosaurs

Parvicursor is a genus of tiny maniraptoran dinosaur with long slender legs for fast running.

<span class="mw-page-title-main">Alvarezsauridae</span> Family of extinct long-legged dinosaurs

Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests.

<i>Aucasaurus</i> Extinct genus of dinosaurs

Aucasaurus is a genus of medium-sized abelisaurid theropod dinosaur from Argentina that lived during the Late Cretaceous of the Anacleto Formation. It was smaller than the related Carnotaurus, although more derived in some ways, such as its extremely reduced arms and almost total lack of fingers. The type skeleton is complete to the thirteenth caudal vertebra, and so is relatively well understood, and is the most complete abelisaurid yet described. However, the skull is damaged, causing some paleontologists to speculate that it was involved in a fight prior to death.

<i>Velocisaurus</i> Extinct genus of dinosaurs

Velocisaurus is a genus of noasaurid theropod dinosaur from the Late Cretaceous period of Argentina.

<i>Mononykus</i> Extinct genus of dinosaurs

Mononykus is a genus of alvarezsaurid dinosaur that lived during the Late Cretaceous in what is now Asia on the Nemegt Formation, about 70 million years ago.

<i>Andesaurus</i> Extinct genus of dinosaurs

Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.

<i>Citipati</i> Genus of oviraptorid dinosaur

Citipati is a genus of oviraptorid dinosaur that lived in Asia during the Late Cretaceous period, about 75 million to 71 million years ago. It is mainly known from the Ukhaa Tolgod locality at the Djadochta Formation, where the first remains were collected during the 1990s. The genus and type species Citipati osmolskae were named and described in 2001. A second species from the adjacent Zamyn Khondt locality may also exist. Citipati is one of the best-known oviraptorids thanks to a number of well-preserved specimens, including individuals found in brooding positions atop nests of eggs, though most of them were initially referred to the related Oviraptor. These nesting specimens have helped to solidify the link between non-avian dinosaurs and birds.

<i>Patagonykus</i> Extinct genus of dinosaurs

Patagonykus is a genus of theropod dinosaur from the Upper Cretaceous of Argentina. This alvarezsauroid was discovered in exposures of the Portezuelo Formation (Turonian-Coniacian) of the Rio Neuquén Subgroup in the Neuquén Basin, Neuquen Province of Patagonia, Argentina. The holotype consists of an incomplete but well-preserved skeleton, lacking a skull, but including many vertebrae, the coracoids, a partial forelimb, pelvic girdle, and hindlimbs. Patagonykus has been classed with the Alvarezsauridae, a family which includes such taxa as the Mongolian Mononykus and the Argentinian Alvarezsaurus. In 2010 Gregory S. Paul estimated its length at 1 meter and its weight at 3.5 kg.

<span class="mw-page-title-main">Anacleto Formation</span> Geologic formation in Argentina

The Anacleto Formation is a geologic formation with outcrops in the Argentine Patagonian provinces of Mendoza, Río Negro, and Neuquén. It is the youngest formation within the Neuquén Group and belongs to the Río Colorado Subgroup. Formerly that subgroup was treated as a formation, and the Anacleto Formation was known as the Anacleto Member.

<span class="mw-page-title-main">Maniraptoriformes</span> Clade of dinosaurs

Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptorans. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of Ornithomimus and birds, and all descendants of that common ancestor."

Amargatitanis is a genus of dicraeosaurid sauropod dinosaur from the Barremian-age La Amarga Formation of Neuquén, Argentina. It is known from a single, incomplete postcranial skeleton consisting of a partial hindlimb, ischium, and two vertebrae. These remains were unearthed by Argentine paleontologist José Bonaparte in 1983 during an expedition by the Museo Argentino de Ciencias Naturales and later described as a new genus and species, Amargatitanis macni by Sebastián Apesteguía. The genus name comes from the words Amarga, where the fossils were collected, and titanis meaning "titan". Its species name is in reference to the MACN, where the remains are stored.

<i>Achillesaurus</i> Extinct genus of dinosaurs

Achillesaurus is a genus of alvarezsaurid theropod dinosaur from the Santonian-age Upper Cretaceous Bajo de la Carpa Formation of Rio Negro, Argentina. It was named in reference to Achilles' heel, because diagnostic features are found there for these animals. The genus was a relatively large, basal alvarezsaurid, and a contemporary of Alvarezsaurus. Achillesaurus is based on MACN-PV-RN 1116, a partial skeleton including a sacral vertebra, four tail vertebrae, part of the left thighbone, shin and foot, and the left ilium. Agustín Martinelli and Ezequiel Vera, who described the specimen, performed a phylogenetic analysis and found their new genus to be an alvarezsaurid with an unresolved relationship to Alvarezsaurus and more derived alvarezsaurids. Makovicky, Apesteguía & Gianechini (2012) argued that Achillesaurus might actually be a junior synonym of Alvarezsaurus which, according to the authors, "is known from the same formation and from which it [i.e. Achillesaurus] differs trivially."

<i>Ceratonykus</i> Extinct genus of dinosaurs

Ceratonykus is a monospecific genus of alvarezsaurid dinosaur from Mongolia that lived during the Late Cretaceous in what is now the Barun Goyot Formation. The type and only species, Ceratonykus oculatus, is known from a fragmentary skeleton, including an incomplete skull, of an adult individual. It was named and described in 2009 by Vladimir Alifanov and Rinchen Barsbold. Its describers questioned the traditional placement of alvarezsaurs in Theropoda, instead suggesting they were ornithischians, but this has not been accepted since. Ceratonykus has an estimated length of 75 centimetres and weight of 760 grams. It has been considered as a possible junior synonym of Parvicursor.

<span class="mw-page-title-main">Alvarezsauroidea</span> Extinct superfamily of dinosaurs

Alvarezsauroidea is a group of small maniraptoran dinosaurs. Alvarezsauroidea, Alvarezsauridae, and Alvarezsauria are named for the historian Gregorio Álvarez, not the more familiar physicist Luis Alvarez, or his son geologist Walter Alvarez who jointly proposed that the Cretaceous–Paleogene extinction event was caused by an impact event. The group was first formally proposed by Choiniere and colleagues in 2010, to contain the family Alvarezsauridae and non-alvarezsaurid alvarezsauroids, such as Haplocheirus, which is the basalmost of the Alvarezsauroidea. The discovery of Haplocheirus extended the stratigraphic evidence for the group Alvarezsauroidea about 63 million years further in the past. The division of Alvarezsauroidea into the Alvarezsauridae and the non-alvarezsaurid alvarezsauroids is based on differences in their morphology, especially in their hand morphology.

<i>Linhenykus</i> Extinct genus of dinosaurs

Linhenykus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous of Inner Mongolia, China. It is the most basal known member of the Parvicursorinae. The genus gets its name from Linhe, a city near the site where the fossil was first found and Greek nykus, "claw". The specific name is derived from Greek monos, "single", and daktylos, "finger", a reference to the fact that it is the only known non-avian dinosaur to have had but a single digit.

<i>Bonapartenykus</i> A large alvarezsauroid dinosaur

Bonapartenykus is a monospecific genus of alvarezsauroid dinosaur from Argentina that lived during the Late Cretaceous (Campanian-Maastrichtian) in what is now the upper Allen Formation of the Río Negro Province. The type and only species, Bonapartenykus ultimus, is known from a nearly articulated but partial skeleton that was found in close association to two incomplete eggs and several clusters of eggshells belonging to the oogenus Arriagadoolithus. Bonapartenykus was named in 2012 by Federico L. Agnolin, Jaime E. Powell, Fernando E. Novas and Martin Kundrát. Bonapartenykus has an estimated length of 2.5 m (8.2 ft) and weight of 72 kg (159 lb), making it the largest member of the clade Alvarezsauroidea.

<i>Overoraptor</i> Extinct genus of theropod dinosaurs

Overoraptor is an extinct genus of paravian theropod of uncertain affinities from the Late Cretaceous Huincul Formation of Argentinian Patagonia. The genus contains a single species, O. chimentoi, known from several bones of the hands, feet, and hips alongside some vertebrae.

Khulsanurus is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Barungoyot Formation of the Khulsan Locality in the Gobi Desert region of Mongolia. The type and only species is Khulsanurus magnificus.

Sidersaura is an extinct genus of rebbachisaurid sauropod dinosaur from the Late Cretaceous Huincul Formation of Argentina. The genus contains a single species, S. marae, known from the remains of four individuals. Sidersaura represents one of the largest known rebbachisaurids.

References

  1. Holtz Jr., Thomas R. (2012). "Holtz's Genus List" (PDF).
  2. Paul, Gregory S. (2010). The Princeton Field Guide to Dinosaurs (1st ed.). Princeton University Press. pp.  126. ISBN   978-0691137209. JSTOR   j.ctt7sv5n.
  3. Bonaparte, José F. (1991). "Los vertebrados fósiles de la formación Río Colorado, de la ciudad de Neuquén y cercanías, Cretácico Superior, Argentina" [The fossil vertebrates of the Colorado River formation, from the city of Neuquén and surroundings, Upper Cretaceous, Argentina]. Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia". Paleontología (in Spanish). 4 (3): 15–123. ISSN   0524-9511.
  4. 1 2 Chiappe, Luis M.; Norell, Mark A.; Clark, James M. (5 December 2002). "The Cretaceous, Short-Armed Alvarezsauridae: Mononykus and Its Kin". In Chiappe, Luis M.; Witmer, Lawrence M. (eds.). Mesozoic Birds: Above the Heads of Dinosaurs. University of California Press (published December 2002). pp. 87–120. ISBN   978-0520200944.
  5. Choiniere, Jonah N.; Xu, Xing; Clark, James M.; Forster, Catherine A.; Guo, Yu; Han, Fenglu (29 January 2010). "A Basal Alvarezsauroid Theropod from the Early Late Jurassic of Xinjiang, China". Science . 327 (5965): 571–574. Bibcode:2010Sci...327..571C. doi:10.1126/science.1182143. PMID   20110503. S2CID   36904501.