Macrodontopteryx Temporal range: Early Eocene | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | † Odontopterygiformes |
Family: | † Pelagornithidae |
Genus: | † Macrodontopteryx Harrison & C.A.Walker, 1976 |
Species: | †M. oweni |
Binomial name | |
†Macrodontopteryx oweni Harrison & C.A.Walker, 1976 | |
Synonyms | |
see text |
Macrodontopteryx is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These animals were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty. [1]
Only a single species, Macrodontopteryx oweni, is known to date. Its remains were found in the Ypresian (Early Eocene) London Clay of the Isle of Sheppey (England), where they were deposited about 50 Ma (million years ago) in the Mammal Paleogene zone MP8-9. The fossils were at first assigned to Argillornis longipennis, which is nowadays recognized as junior synonym of Dasornis emuinus , a larger pseudotooth bird than M. oweni. Due to the scarcity of their fossil remains, is not clear whether the pseudotooth bird species described as Pseudodontornis longidentata is not the same as M. oweni, though its size suggests that it is more likely to be synonymous with D. emuinus. In any case, Macrodontopteryx itself may well be no distinct genus but rather a large member of the contemporary and sympatric Odontopteryx , or a young or small Dasornis. [2]
The only known specimen (BMNH A1) is an incomplete and rather crushed skull with much all of the beak missing. In the living animal it would have measured roughly 30–35 cm (one foot). The wingspan was probably around 4 m (13 ft). [3]
If Macrodontopteryx is indeed a distinct genus, perhaps some pseudotooth bird fossils found in Lutetian (Middle Eocene, MP11-13, about 45 Ma) deposits at Etterbeek (Belgium) belong therein. Like the holotype BMNH A1, they have been assigned to Argillornis longipennis in the past; at least some of the material [4] placed in that taxon at one time or another – but not its syntype humerus pieces – seems too small to be of an adult Dasornis. [5]
An Early Oligocene (Stampian, MP21-23, about 32 Ma) distal radius from Hamstead, Isle of Wight (England) was also assigned to Macrodontopteryx. It is without doubt from a very large and flight-capable bird, but little else can be said about it. An equally old cervical vertebra fragment (specimen BMNH A-4413) and a toe phalanx from nearby Yarmouth were, mainly due to their size, believed to be related to the enigmatic large flightless bird (and presumed ratite) Eleutherornis . They were described as Proceriavis martini , [6] but have never been compared in detail to pseudotooth bird fossils. It cannot be ruled out that Proceriavis belongs in the Pelagornithidae, but whether it would be a junior synonym of Macrodontopteryx is highly uncertain given the 20-million-year difference between the Isle of Sheppey and Isle of Wight material. [7] Recent analysis suggests that the material referred to as Proceriavis is in fact ratite in nature. [8]
Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae, Anseranatidae, and Anatidae, the largest family, which includes over 170 species of waterfowl, among them the ducks, geese, and swans. Most modern species in the order are highly adapted for an aquatic existence at the water surface. With the exception of screamers, males have penises, a trait that has been lost in the Neoaves. Due to their aquatic nature, most species are web-footed.
Struthionidae is a family of flightless birds, containing the extant ostriches and their extinct relatives. The two extant species of ostrich are the common ostrich and Somali ostrich, both in the genus Struthio, which also contains several species known from Holocene fossils such as the Asian ostrich. The common ostrich is the more widespread of the two living species, and is the largest living bird species. The extinct genus Pachystruthio from the Late Pliocene-Early Pleistocene of Eurasia is one of the largest birds ever.
Traditionally, the bird order Apodiformes contained three living families: the swifts (Apodidae), the treeswifts (Hemiprocnidae), and the hummingbirds (Trochilidae). In the Sibley-Ahlquist taxonomy, this order is raised to a superorder Apodimorphae in which hummingbirds are separated as a new order, Trochiliformes. With nearly 450 species identified to date, they are the most diverse order of birds after the passerines.
The mousebirds are birds in the order Coliiformes. They are the sister group to the clade Cavitaves, which includes the Leptosomiformes, Trogoniformes (trogons), Bucerotiformes, Piciformes and Coraciformes. This group is now confined to sub-Saharan Africa, and it is the only bird order confined entirely to that continent, with the possible exception of turacos which are considered by some as the distinct order Musophagiformes, and the cuckoo roller, which is the only member of the order Leptosomiformes, and which is found in Madagascar but not mainland Africa. Mousebirds had a wider range in the Paleogene, with a widespread distribution in Europe and North America during the Paleocene.
Elopteryx is a genus of, perhaps troodontid, maniraptoran theropod dinosaur based on fragmentary fossils found in late Cretaceous Period rocks of Romania. The single species, Elopteryx nopcsai, is known only from very incomplete material, and therefore is considered a nomen dubium by most paleontologists.
Osteodontornis is an extinct seabird genus. It contains a single named species, Osteodontornis orri, which was described quite exactly one century after the first species of the Pelagornithidae was. O. orri was named after Santa Barbara Museum of Natural History paleontologist Phil C. Orr, for his recognition of the importance of the specimen.
Presbyornis is an extinct genus of anseriform bird. It contains two unequivocally accepted species; the well-known P. pervetus and the much lesser-known P. isoni. P. pervetus was approximately the size and shape of a goose, but with longer legs; P. isoni, known from a few bones, was much larger, more than swan-sized. Other fossils, more doubtfully assigned to this genus, are also known.
The Pelagornithidae, commonly called pelagornithids, pseudodontorns, bony-toothed birds, false-toothed birds or pseudotooth birds, are a prehistoric family of large seabirds. Their fossil remains have been found all over the world in rocks dating between the Early Paleocene and the Pliocene-Pleistocene boundary.
Diogenornis is an extinct genus of ratites, that lived during the Early Eocene. It was described in 1983 by Brazilian scientist Herculano Marcos Ferraz de Alvarenga based on fossils found in the Itaboraí Formation in southeastern Brazil. The type species is D. fragilis. While initially considered a member of the family Opisthodactylidae, further examination of the fossil remains showed that it was more similar to the modern rhea. According to Gerald Mayr, Diogenornis is best considered a stem-group member of the Rheidae. It grew to about two thirds the size of the modern greater rhea, at about 90 centimetres (3.0 ft) of height.
Odontopteryx is a genus of the extinct pseudotooth birds or pelagornithids. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Caspiodontornis is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Cyphornis is a genus of the prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Dasornis is a genus of prehistoric pseudotooth birds. These were probably close relatives of either pelicans and storks or waterfowl; they are placed in the order Odontopterygiformes to account for this uncertainty.
Gigantornis eaglesomei is a very large prehistoric bird described from a fragmentary specimen from the Eocene of Nigeria. It was originally described as a representative of the albatross family, Diomedeidae, but was later referred to the pseudotooth or bony-toothed bird family, Pelagornithidae. Gigantornis is one of the largest pseudotooth birds, with an inferred wingspan of about 6 m (20 ft) it is among the largest birds ever.
Palaeochenoides is a genus of the prehistoric pseudotooth birds of somewhat doubtful validity. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Pelagornis is a widespread genus of prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or waterfowl, and are placed here in the order Odontopterygiformes to account for this uncertainty.
Pseudodontornis is a rather disputed genus of the prehistoric pseudotooth birds. The pseudotooth birds or pelagornithids were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty. Up to five species are commonly recognized in this genus.
Tympanonesiotes is a somewhat doubtfully valid genus of the prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertainty.
Lithornis is a genus of extinct paleognathous birds. Although Lithornis was able to fly well, their closest relatives are the extant tinamous and ratites.
Geranoididae is a clade of extinct birds from the early to late Eocene and possibly early Oligocene of North America and Europe. These were mid-sized, long-legged flightless birds. Recent research shows that these birds may actually be palaeognaths related to ostriches.