Ambopteryx Temporal range: Late Jurassic, | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Family: | † Scansoriopterygidae |
Genus: | † Ambopteryx Wang et al., 2019 |
Species: | †A. longibrachium |
Binomial name | |
†Ambopteryx longibrachium Wang et al., 2019 | |
Ambopteryx (meaning "both wing") is a genus of scansoriopterygid dinosaur from the Oxfordian stage of the Late Jurassic of China. It is the second dinosaur to be found with both feathers and bat-like membranous wings. Yi, the first such dinosaur, was described in 2015 and is the sister taxon to Ambopteryx. The holotype specimen is thought to be a sub-adult or adult. The specimen is estimated to have had a body length of 32 centimetres (13 in) and a weight of 306 grams (10.8 oz). The genus includes one species, Ambopteryx longibrachium. [1]
The generic name Ambopteryx is derived from the Latin word ambo meaning "both" and Ancient Greek word πτέρυξ (pteryx) meaning "wing", a reference to the animal's membranous wings and bird-like body plan. The specific name longibrachium is derived from the Latin words longus meaning "long" and brachium meaning "upper arm". [1]
The holotype specimen, IVPP V24192, is an articulated and nearly complete skeleton with associated soft tissue preserved on a slab and a counter slab. It was recovered in 2017 from the stratigraphic equivalent of the Haifanggou Formation near Wubaiding Village in the Liaoning Province and belongs to the Yanliao Biota. [1]
The type specimen is interpreted as either a subadult or an adult based on the fusion of the pygostyle. Its wing membrane is preserved as a continuous brownish layer on the matrix surrounding the left hand, right forelimb, and abdomen. The head, neck, and shoulder regions are covered with dense layers of feather. [1]
Like other scansoriopterygids, Ambopteryx had a short, blunt head and a hyper-elongate third finger. It was previously believed that scansoriopterygids used their third finger to extract grubs from wood, like the extant aye-aye. However, Wang et al. think the primary function of the third finger was the attachment of the wing membrane, or patagium, as their fingers were probably surrounded by membranous tissue and had limited mobility. [1]
Among scansoriopterygids, Ambopteryx was most closely related to Yi, with which it shared a "styliform element", an elongate bone that extended from the distal (far) end of the ulna. The styliform element was slightly curved and tapered towards the distal end. It supported the wing membrane, which stretched from the third finger down to the abdomen. [1]
The forelimbs of Ambopteryx were about 1.3 times longer than the hindlimbs, with the ulna being shorter than the humerus and almost twice as wide as the width of the radius. Unlike most other non-avian theropods, it had a short tail that lacked a transitional point and ended in a pygostyle, a set of fused tail vertebrae. The only other non-avian theropod that are known to have had a pygostyle are certain oviraptorosaurs and therizinosaurs. [1]
The abdominal region of Ambopteryx contains a small number of gastroliths and large fragments of what appears to be bone. The bone probably represents stomach contents. Previously, the diet of scansoriopterygids was unknown but their unusual dental morphology and the presence of gastroliths and possible bone fragments in Ambopteryx suggests they were omnivorous. [1]
Theropoda, whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago (Ma) and included the majority of large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species.
Protarchaeopteryx is a genus of turkey-sized feathered theropod dinosaur from China. Known from the Jianshangou bed of the Yixian Formation, it lived during the early Aptian age of the Early Cretaceous, approximately 124.6 million years ago. It was probably a herbivore or omnivore, although its hands were very similar to those of small carnivorous dinosaurs. It appears to be one of the most basal members of the Oviraptorosauria, closely related to Incisivosaurus, or a taxon slightly less closely related to birds than oviraptorosaurs were.
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, Oviraptorosauria, and Therizinosauria. Ornitholestes and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period, and survive today as living birds.
Beipiaosaurus is a genus of therizinosauroid theropod dinosaurs that lived in Asia during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the discovery of Yutyrannus, Beipiaosaurus were among the heaviest dinosaurs known from direct evidence to be feathered. Beipiaosaurus is known from three reported specimens. Numerous impressions of feather structures were preserved that allowed researchers to determine the feathering color which turned out to be brownish.
Deinocheirus is a genus of large ornithomimosaur that lived during the Late Cretaceous around 70 million years ago. In 1965, a pair of large arms, shoulder girdles, and a few other bones of a new dinosaur were first discovered in the Nemegt Formation of Mongolia. In 1970, this specimen became the holotype of the only species within the genus, Deinocheirus mirificus; the genus name is Greek for "horrible hand". No further remains were discovered for almost fifty years, and its nature remained a mystery. Two more complete specimens were described in 2014, which shed light on many aspects of the animal. Parts of these new specimens had been looted from Mongolia some years before, but were repatriated in 2014.
The patagium is a membranous body part that assists an animal in obtaining lift when gliding or flying. The structure is found in extant and extinct groups of flying and gliding animals including bats, birds, some dromaeosaurs, pterosaurs, gliding mammals, some flying lizards, and flying frogs. The patagium that stretches between an animal's hind limbs is called the uropatagium or the interfemoral membrane.
Anurognathus is an extinct genus of small pterosaur from the Late Jurassic Altmühltal Formation of Germany.
Scansoriopteryx is a genus of maniraptoran dinosaur. Described from only a single juvenile fossil specimen found in Liaoning, China, Scansoriopteryx is a sparrow-sized animal that shows adaptations in the foot indicating an arboreal (tree-dwelling) lifestyle. It possessed an unusual, elongated third finger which may have supported a membranous wing, much like the related Yi qi. The type specimen of Scansoriopteryx also contains the fossilized impression of feathers.
Scansoriopterygidae is an extinct family of climbing and gliding maniraptoran dinosaurs. Scansoriopterygids are known from five well-preserved fossils, representing four species, unearthed in the Tiaojishan Formation fossil beds of Liaoning and Hebei, China.
Sinocalliopteryx is a genus of carnivorous compsognathid theropod dinosaurs from the Lower Cretaceous Yixian Formation of China.
Sinornis is a genus of enantiornithean birds from the Lower Cretaceous Jiufotang Formation of the People's Republic of China.
Archaeorhynchus is a genus of beaked avialan stem-birds from the early Cretaceous period. A fossil of its only known species, Archaeorhynchus spathula, was first reported in 2005 by Zhou & Zhang to have been found in Yixian Formation rocks at Yixian, Liaoning province, China, showing a well-preserved and essentially complete skeleton. Two more complete specimens were found in Lower Cretaceous deposits of Jianchang, Liaoning, northeastern China, preserving new anatomical information. These deposits are 120 million years old, whereas the original specimen was 125 million years old, meaning the age range for this species is 125-120Ma.
Zhongornis is a genus of primitive maniraptoran dinosaurs that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008.
Limusaurus is a genus of theropod dinosaur that lived in what is now China during the Late Jurassic, around 161 to 157 million years ago. The type and only species Limusaurus inextricabilis was described in 2009 from specimens found in the Upper Shishugou Formation in the Junggar Basin of China. The genus name consists of the Latin words for "mud" and "lizard", and the species name means "impossible to extricate", both referring to these specimens possibly dying after being mired. Limusaurus was a small, slender animal, about 1.7 m in length and 15 kg (33 lb) in weight, which had a long neck and legs but very small forelimbs. It underwent a drastic morphological transformation as it aged: while juveniles were toothed, these teeth were completely lost and replaced by a beak with age. Several of these features were convergently similar to the later ornithomimid theropods as well as the earlier non-dinosaurian shuvosaurids.
Haplocheirus is an extinct genus of theropod dinosaur from the Middle Jurassic Shishugou Formation of Xinjiang in China. It is generally considered to be an alvarezsauroid, although some researchers have questioned this assignment. The genus contains a single species, H. sollers, which is known from a mostly complete skeleton including the skull.
Martharaptor is a genus of therizinosauroid theropod dinosaurs from the Early Cretaceous of the Cedar Mountain Formation in Utah. They can be distinguished from other therizinosauroids by means of several features of the skeleton which were intermediate between early therizinosaurs such as Falcarius and Beipiaosaurus, and more "advanced" members of the group like therizinosaurids. The deep and homogeneous hand claws clearly differ from the case in early therizinosauroids, but the foot has not yet acquired the robust morphology of therizinosaurids.
Yi is a genus of scansoriopterygid dinosaurs from the Late Jurassic of China. Its only species, Yi qi, is known from a single fossil specimen of an adult individual found in Middle or Late Jurassic Tiaojishan Formation of Hebei, China, approximately 159 million years ago. It was a small, possibly tree-dwelling (arboreal) animal. Like other scansoriopterygids, Yi possessed an unusual, elongated third finger, that appears to have helped to support a membranous gliding plane made of skin. The planes of Yi qi were also supported by a long, bony strut attached to the wrist. This modified wrist bone and membrane-based plane is unique among all known dinosaurs, and might have resulted in wings similar in appearance to those of bats.
Zhongjianosaurus is a genus of dromaeosaurid belonging to the Microraptoria. Believed to hail from the Yixian Formation, specifically the middle of the Jehol Biota, it is the smallest known microraptorine thus far discovered and one of the smallest non-avian theropod dinosaurs.
Serikornis is a genus of small, feathered anchiornithid dinosaur from the Upper Jurassic Tiaojishan Formation of Liaoning, China. It is represented by the type species Serikornis sungei.
Caihong is a genus of small paravian theropod dinosaur from China that lived during the Late Jurassic period.