Zhongornis Temporal range: Early Cretaceous, | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | Avialae |
Genus: | † Zhongornis Gao et al., 2008 |
Species: | †Z. haoae |
Binomial name | |
†Zhongornis haoae Gao et al., 2008 | |
Zhongornis (meaning "intermediate bird" [1] ) is a genus of primitive maniraptoran dinosaurs that lived during the Early Cretaceous. It was found in rocks of the Yixian Formation in Lingyuan City (China), and described by Gao et al. in 2008. [2]
Zhongornis has only one described species, Zhongornis haoae. The only specimen is a fossil slab and counterslab numbered D2455/6. It is in the collection of the Dalian Natural History Museum. It is a fairly complete skeleton about eight centimeters in length. Pores in the bones and unfused sutures in the skeleton indicate that the specimen was a juvenile, but the authors believe that it was developed enough to erect a new taxon on the basis of its unique morphological characters. There are feather impressions preserved on the right hand and also probable tail feathers preserved near the left foot. Zhongornis had a beaked mouth with no teeth. The tail is proportionately short, has thirteen vertebrae, and no pygostyle. The third finger has only two phalangeal bones, unlike non-avian dinosaurs and Confuciusornis , and more like Enantiornithes and more advanced avialans. These features and a cladistic analysis indicate that Zhongornis is the sister group to all pygostylia, meaning that it is intermediate between long-tailed Avialae, like Archaeopteryx , and more advanced taxa, like Confuciusornis.
Zhongornis provides important anatomical information about the evolutionary transition from primitive basal Avialae like Archaeopteryx, which had a long bony tail and a dinosaur-like third finger, to the more advanced avialans like the Enantiornithes, which had reduced third fingers and tails fused into rigid pygostyles. Zhonghornis is the only fossil ever found that seems to be intermediate in these features. It appears to have one less bone in the third finger than Archaeopteryx, and one more than Longipteryx , suggesting that it is an intermediate between the two. [2] Zhongornis also seems to be intermediate in its tail anatomy. It has only thirteen caudal vertebrae, far less than the 22 in Archaeopteryx. None of the vertebral centra are fused, but the last four do form a continuous lateral flange, implying that this specimen had an incipient pygostyle. Previous to this fossil Sanz et al. (1992) suggested that the evolution of the pygostyle may have proceeded as the numerous vertebrae of the tail became very small and highly ankylosed. [3] Zhongornis suggests that shortening of the tail, and a large reduction in the number of vertebrae, preceded the origin of the pygostyle in the evolution of at least one bird language.
After a detailed restudy, O'Connor and Sullivan suggest that this juvenile bird from the Early Cretaceous Jehol fauna of China actually possesses 20, rather than 14, tail vertebrae; further, this tail is very similar to those of Epidexipteryx (a scansoriopterygid) and Caudipteryx (an oviraptorosaur). Based on this and other features, the authors reinterpreted Zhongornis as the sister taxon of scansoriopterygids, and further suggested that this clade (Zhongornis + Scansoriopterygidae) is the sister group of Oviraptorosauria. [4] Other phylogenetic analyses, including a larger dataset, don't consider the hypothesis that Zhongornis is a scansoriopterygid, instead finding it in its traditional position as a primitive relative of pygostylians, or as a close relative of Confuciusornis. [5]
Caudipteryx is a genus of small oviraptorosaur dinosaurs that lived in Asia during the Early Cretaceous, around 124.6 million years ago. They were feathered and extremely birdlike in their overall appearance, to the point that some paleontologists suggested it was a bird. Two species have been described: C. zoui, in 1998, and C. dongi, in 2000.
Confuciusornis is a genus of basal crow-sized avialan from the Early Cretaceous Period of the Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, Confuciusornis had a toothless beak, but closer and later relatives of modern birds such as Hesperornis and Ichthyornis were toothed, indicating that the loss of teeth occurred convergently in Confuciusornis and living birds. It was thought to be the oldest known bird to have a beak, though this title now belongs to an earlier relative Eoconfuciusornis. It was named after the Chinese moral philosopher Confucius. Confuciusornis is one of the most abundant vertebrates found in the Yixian Formation, and several hundred complete specimens have been found.
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to Ornithomimus velox. It contains the major subgroups Avialae, Dromaeosauridae, Troodontidae, Oviraptorosauria, and Therizinosauria. Ornitholestes and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period, and survive today as living birds.
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.
Beipiaosaurus is a genus of therizinosauroid theropod dinosaurs that lived in Asia during the Early Cretaceous in the Yixian Formation. The first remains were found in 1996 and formally described in 1999. Before the discovery of Yutyrannus, Beipiaosaurus were among the heaviest dinosaurs known from direct evidence to be feathered. Beipiaosaurus is known from three reported specimens. Numerous impressions of feather structures were preserved that allowed researchers to determine the feathering color which turned out to be brownish.
Oviraptorosaurs are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from Caudipteryx, which was the size of a turkey, to the 8-meter-long, 1.4-ton Gigantoraptor. The group is close to the ancestry of birds. Some researchers such as Maryanska et al (2002) and Osmólska et al. (2004) have proposed that they may represent primitive flightless birds. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.
Avimimus, meaning "bird mimic", is a genus of oviraptorosaurian theropod dinosaur, named for its bird-like characteristics, that lived in the late Cretaceous in what is now Mongolia, around 85 to 70 million years ago.
Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main component of the uropygium, a structure colloquially known as the bishop's nose, parson's nose, pope's nose, or sultan's nose. This is the fleshy protuberance visible at the posterior end of a bird that has been dressed for cooking. It has a swollen appearance because it also contains the uropygial gland that produces preen oil.
Scansoriopteryx is a genus of maniraptoran dinosaur. Described from only a single juvenile fossil specimen found in Liaoning, China, Scansoriopteryx is a sparrow-sized animal that shows adaptations in the foot indicating an arboreal (tree-dwelling) lifestyle. It possessed an unusual, elongated third finger which may have supported a membranous wing, much like the related Yi qi. The type specimen of Scansoriopteryx also contains the fossilized impression of feathers.
Scansoriopterygidae is an extinct family of climbing and gliding maniraptoran dinosaurs. Scansoriopterygids are known from five well-preserved fossils, representing four species, unearthed in the Tiaojishan Formation fossil beds of Liaoning and Hebei, China.
Jeholornis is a genus of avialan dinosaurs that lived between approximately 122 and 120 million years ago during the early Cretaceous Period in China. Fossil Jeholornis were first discovered in the Jiufotang Formation in Hebei Province, China and additional specimens have been found in the older Yixian Formation.
Archaeopterygidae is a group of paravian dinosaurs, known from the latest Jurassic and earliest Cretaceous of Europe. In most current classifications, it contains only the genera Archaeopteryx and Wellnhoferia. As its name suggests, Protarchaeopteryx was also once referred to this group, but most paleontologists now consider it an oviraptorosaur. Other referred genera, like Jurapteryx, Wellnhoferia, and "Proornis", are probably synonymous with Archaeopteryx or do not belong into this group. Jinfengopteryx was originally described as an archaeopterygid, though it was later shown to be a troodontid. A few studies have recovered Anchiornis and Xiaotingia to also be members of the Archaeopterygidae, though most subsequent analyses have failed to arrive at the same result. Uncertainties still exist, however, and it may not be possible to confidently state whether archaeopterygids are more closely related to modern birds or to deinonychosaurs barring new and better specimens of relevant species. Teeth attributable to archaeopterygids are known from the earliest Cretaceous (Berriasian) Cherves-de-Cognac locality and the Angeac-Charente bonebed of France.
Sapeornis is a monotypic genus of avialan dinosaurs which lived during the early Cretaceous period. Sapeornis contains only one species, Sapeornis chaoyangensis.
Avialae is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used.
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their closest extinct relatives. Confuciusornithidae contains four genera, possessing both shafted and non-shafted (downy) feathers. They are also noted for their distinctive pair of ribbon-like tail feathers of disputed function.
Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avialans, which include diverse extinct taxa as well as the over 10,000 species of living birds. Basal members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. A number of differing scientific interpretations of the relationships between paravian taxa exist. New fossil discoveries and analyses make the classification of Paraves an active subject of research.
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces.
Epidexipteryx is a genus of small paravian dinosaurs, known from one fossil specimen in the collection of the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. Epidexipteryx represents the earliest known example of ornamental feathers in the fossil record.
Ambopteryx is a genus of scansoriopterygid dinosaur from the Oxfordian stage of the Late Jurassic of China. It is the second dinosaur to be found with both feathers and bat-like membranous wings. Yi, the first such dinosaur, was described in 2015 and is the sister taxon to Ambopteryx. The holotype specimen is thought to be a sub-adult or adult. The specimen is estimated to have had a body length of 32 centimetres (13 in) and a weight of 306 grams (10.8 oz). The genus includes one species, Ambopteryx longibrachium.